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Copyright ©1999 by The Resilience Alliance*

Rey Benayas, J. M., S. M. Scheiner, M. García Sánchez-Colomer, and C. Levassor. 1999. Commonness and rarity: theory and application of a new model to Mediterranean montane grasslands. Conservation Ecology [online] 3(1): 5. Available from the Internet. URL: http://www.consecol.org/vol3/iss1/art5/


A version of this article in which text, figures, tables, and appendices are separate files may be found by following this link.

Research

Commonness and Rarity: Theory and Application of a New Model to Mediterranean Montane Grasslands

José M. Rey Benayas1, Samuel M. Scheiner2, Manuel García Sánchez-Colomer3, and Catherine Levassor4


1Universidad de Alcalá; 2Arizona State University West; 3CEDEX; 4Universidad Autónoma de Madrid




ABSTRACT

We examined patterns of commonness and rarity among plant species in montane wet grasslands of Iberia. This examination is set within two contexts. First, we expanded on an earlier scheme for classifying species as common or rare by adding a fourth criterion, the ability of that species to occupy a larger or smaller fraction of its potential suitable habitats, i.e., habitat occupancy. Second, we explicated two theories, the superior organism theory and the generalist/specialist trade-off theory. The data consisted of 232 species distributed among 92 plots. The species were measured for mean local abundance, size of environmental volume occupied, percentage of volume occupied, range within Iberia, and range in Europe and the Mediterranean basin. In general, all measures were positively correlated, in agreement with the superior organism theory. However, specialist species were also found. Thus, patterns of commonness and rarity may be due to a combination of mechanisms. Analyses such as ours can also be used as a first step in identifying habitats and species that may be endangered.

KEY WORDS: commonness, endangered species, generalist/specialist trade-off, geographic range, habitat occupancy, habitat specificity, Iberia, local abundance, montane grasslands, rarity, superior organism theory.

Published March 5, 1999.


INTRODUCTION

Why are some species common and others rare? This question is at the heart of much ecological research and has reached special prominence with current concerns over species conservation and global change (Kunin and Gaston 1997). In this paper, we examine patterns of commonness and rarity among plant species in montane wet grasslands at local, regional, and continental scales. We propose a new conceptual model and apply it to field data. In doing so, we expand on current ideas about commonness and rarity and examine the predictions of two theories.

A touchstone for current discussions of commonness and rarity is the scheme by Rabinowitz (1981), who pointed out that there are actually many forms of rarity (Table 1A). She classified species based on three criteria: geographic range (wide or narrow), habitat specificity (broad or restricted), and local abundance (somewhere large or everywhere small). Only one of the eight possible combinations (wide range, broad habitat specificity, and somewhere large local abundance) is classified as common. The other seven each include some form of rarity, and some of them are even questioned to exist. For an in-depth discussion of types of rarity, see Gaston (1994).


Table 1. A scheme for describing types of commonness and rarity.

A) The original proposal by Rabinowitz (1981).

Geographic range Wide Narrow
Habitat specificity Broad Restricted Broad Restricted
Abundance somewhere large Common Predictable Unlikely Endemics
Abundance everywhere small Sparse Non-existent

B) Our new scheme adding the notion of habitat occupancy.

Geographic range Wide Narrow
Habitat specificity Broad Restricted Broad Restricted
Abundance Large Small Large Small Large Small Large Small
Habitat occupancy high Common Widespread Indicator Locally common Non-existent Endemic indicator
Habitat occupancy low Highly dispersed Sparse Locally endangered Potentially endangered Endangered


There is yet a fourth criterion for defining commonness vs. rarity. Within the habitat specificity of a species is the ability of that species to occupy a larger or smaller fraction of its potential suitable habitats, i.e., habitat occupancy. This criterion is theoretically independent of the other three (Table 1B). As far as we are aware, no study has simultaneously examined the relationship among all four criteria. For example, Agrostis castellana is found in a range of mesic forests and grasslands including those of our study. Yet, within Quercus pyrenaica forests in the same area, this species was found in only two of 21 plots (Fernández González 1991). In contrast, Carex demissa is a specialist of wet grasslands and was found within 63% of those habitats in our study (see Results). The distinction between habitat occupancy and habitat specificity is somewhat akin to the distinction between realized and fundamental niche (Hutchinson 1957). We emphasize, though, that this is just a rough analogy, because habitat specificity is measured as the observed distribution of a species. Although the notion of occupancy has been raised previously (e.g., Orians 1997), ours is the first study to analyze its relationship to all of these other components of rarity (however, see Burgman 1989).

For convenience, we modify and expand Rabinowitz's (1981) earlier scheme. Now, only one of the 16 combinations of species characteristics (Table 1B) would be considered common: wide geographic range, broad habitat specificity, large local abundance, and frequent habitat occupation. According to Rabinowitz, the other categories are all forms of rarity. We disagree somewhat by recognizing a category of locally common species, similar to common species but having a narrow geographic range. Species in the categories that we designate as widespread and highly dispersed could also reasonably be considered common. These quibbles, however, do not obviate Rabinowitz's central point that commonness and rarity are complex phenomena. Of course, nature does not fit into neat categories. This binary classification scheme actually stands for a continuum of species types.

The use of wet grasslands is particularly appropriate in this context. These habitats differ from the surrounding land because groundwater seeps result in a moister environment, changes in soil features, and a distinctive plant community. These differences are most conspicuous in the dry season, when patches of green, wet-meadow vegetation, dominated by Juncus, Carex, and hygrophyte graminoid species, stand out against a yellow-brown matrix of senescent annual grasses.

These habitats are rare throughout the Mediterranean region, where they represent important islands of biodiversity (Bernáldez et al. 1993, Rey Benayas et al. 1998). In the mountains of central Spain, they occupy only 9% of the total land area. However, these habitats are much richer in species than are the surrounding plant communities, holding 43.3% of the perennial species in these systems because of the presence of outcropping water in the summer (García Sánchez-Colomer 1998). Also, their species richness per unit area is greater (24.9 species/100 m2 vs. 16.4 species/100 m2; Rey Benayas et al. 1998). Understanding their species composition and why and how they act within the overall context of species' ranges and habitat spectra will help us to design and manage other sorts of diversity islands, either natural or man-made, by fragmentation of former continuous habitats.

Theories

If we are to be successful both in understanding commonness and rarity and in using that knowledge, we must be able to place our information in a theoretical context. Theories are necessary to organize our data and to guide future actions. Currently, we have few general theories about species richness, commonness, and rarity. Most of what go by the name of theory are merely empirical generalizations. Some seem to hold up well to scrutiny (e.g., island biogeography hypotheses; McGuiness 1984, Myers and Giller 1988), whereas others have been called into question (e.g., Rapoport's rule; Rapoport 1982, Stevens 1989, Pagel et al. 1991, Hughes et al. 1996). We explore two theories that purport to explain patterns of commonness and rarity, testing the predictions of each. Both theories have the virtue of starting from the biology of the organism and deriving ecological patterns. Thus, they are mechanistic models rather than simply empirical generalizations.

The first theory is that of Brown (Brown 1984, Brown et al. 1995), which we term the superior organism theory. He begins with the premise that species vary in their abilities to exploit nature. Some species have large fundamental niches, whereas others have narrow ones. Next, he proposes that the center of a species' range is the region where it can exploit the widest range of resource combinations and, therefore, habitats. Here, the species will be at high abundance. As the distance from this center increases, because environmental variables are autocorrelated, the conditions favoring the species become rarer. Its local habitat specificity narrows and its abundance decreases. Eventually, the edge of the geographic range is reached. Those species that begin with a broad niche will have a large geographic range, whereas those with narrow niches will have a small geographic range.

The second theory is the older notion of a generalist/specialist trade-off (e.g., Fox and Morrow 1981, Futuyuma and Moreno 1988). Again, species vary from having broad to narrow niches. However, this habitat specificity exists throughout the geographic range of a species, rather than narrowing toward the edge of the range. Moreover, habitat specificity trades off with local abundance. Where they are found, specialists have high abundances and generalists have lower abundances.

These two theories make alternative predictions concerning patterns of commonness and rarity (Table 2). In its most extreme form, the superior organism theory predicts the existence of two categories: (1) superior species with wide geographic ranges, broad habitat specificity, large local abundances, and high occupancies; and (2) inferior species with the opposite combination. In Table 1B, these categories are termed common and endangered. The trade-off theory also predicts two categories: (1) generalist species with wide geographic ranges, broad habitat specificity, small abundances, and low occupancies; and (2) specialist species with the opposite combination. In Table 1B, these categories are termed sparse and endemic indicators. Of course, both theories allow for intermediate types, but not for all possible combinations of species traits. For example, neither theory predicts the existence of species with narrow geographic ranges and broad habitat specificities, which we termed locally common. For correlations among traits, the superior organism theory predicts positive correlations between geographic range, habitat specificity, local abundance, and habitat occupancy (Gaston and Lawton 1990, Lawton et al. 1994). In contrast, the trade-off theory predicts positive correlations between geographic range and habitat specificity, positive correlations between local abundance and habitat occupancy, and negative correlations between the first group and the second group.


Table 2. Predictions of the superior organism and trade-off theories. Those predictions upheld by the data are shown in boldface. Ecological breadth groups: WGS, wet-grassland specialists; WGG, wet-grassland generalists; MHG, mesic-habitat generalists.

 
Theory 
Superior organism  Trade-off 
Most frequent categories  Common and endangered  Sparse and endemic indicators 
A) Correlations among traits 
   Geographic range and local abundance  positive  negative 
   Habitat specificity and local abundance  positive  negative 
   Habitat specificity and habitat occupancy  positive  negative 
B) Ordering of ecological breadth groups 
   Geographic range  MHG>WGG>WGS  MHG>WGG>WGS 
   Habitat specificity  WGS=WGG>MHG  MHG>WGG>WGS 
   Local abundance  WGS=WGG>MHG  WGS>WGG>MHG 
   Habitat occupancy  MHG>WGG>WGS  WGS>WGG>MHG 
 


An alternative way to consider these local distributions is by first classifying the species based on their species-wide habitat specificity (i.e., ecological breadth). For example, we can classify the wet-grassland species as (1) those that are narrowly specialized to just part of these habitats; (2) those that are found across all types of wet grasslands, but not beyond them; and (3) those that are found across a wide range of mesic habitats, including wet grasslands. Both theories allow for such an array of species. However, the theories make alternative predictions about values of geographic range, habitat specificity, local abundance, and habitat occupancy across these groups (Table 2). The superior organism model predicts that the third group of species will have the greatest geographic range and the highest occupancy. The second and first groups will have progressively lower values for these traits. Habitat specificity within this circumscribed set will be equally high for the first and second groups and lower for the third group. Finally, local abundance will be high for the first and second groups because these habitats are central for these species. In contrast, the trade-off model predicts that local abundance and habitat occupancy will be highest for the first group and progressively lower for the second and third groups. For habitat specificity, it predicts that the third group will be highest, and progressively smaller for the second and first group. Finally, it makes the same prediction as the other theory with respect to the relative rankings of geographic range.


MATERIALS AND METHODS

The study sites consisted of 66 wet-grassland habitats spread among 2069 km2 in the Sierra de Guadarrama of central Spain (40º N and 4º W; Figs. 1 and 2). The climate is mediterranean. Altitude ranges from 560 to 2430 m, mean annual temperature ranges between 12.4 and 6.3 ºC, and total annual precipitation ranges between 456 and 1331 mm. The parent material is mostly granite. We used panchromatic aerial photographs to map the network of wet-grassland habitats in the study area. We selected recent 1:20000-scale stereopairs to generate an updated, accurate map of these habitats. Within them we placed 92 100-m2 plots. Plot locations were chosen to maximize the environmental range sampled. They were placed after field inspections so that larger grasslands with more than one habitat zone (based on visual inspection) received more plots. Most of the wet grasslands (66.7%) received one plot, with a maximum of four plots in one grassland. For small grasslands with only one obvious habitat zone, the plot was placed in the center. For larger grasslands, plots were placed in the approximate center of each habitat zone.


Fig. 1. Map of the study area and location of sample plots in wet grasslands of the Sierra de Guadarrama, Spain.

GIF image file (11 K)


Fig. 2. A sedge-rush habitat in La Salabrosa, Madrid, dominated by Scirpus holoschoenus. The photograph shows the contrast between this wet habitat, due to groundwater seepage, and the surrounding habitats dominated by evergreen oaks (Quercus ilex subsp. rotundifolia ) and xerophytic annuals.

GIF image file (116 K)


We surveyed all perennial vascular plants occurring in the sample plots (Appendices 1 and 2). Over 90% of species in these habitats are perennials. A total of 237 taxa were found, of which 232 could be identified to species. The four species characteristics of local abundance, habitat specificity, habitat occupancy, and geographic range were measured as follows. (See Table 3 for definitions of all measures.) Abundance was estimated by eye using a six-point Braun-Blanquet scale (0, 0.1 = <1% cover, 1 = 1%-10% cover, 2 = 10%-25% cover, 3 = 25%-50% cover, 4 = 50%-75% cover, 5 = 75%-100% cover). Our measure of local abundance was the mean cover value for all plots within which the species appeared. Although Rabinowitz's scheme can be construed to refer to maximal abundance, the mean and the maximum were highly correlated for our data (rS = 0.88, P < 0.0001, n = 232).


Table 3. Definitions of the ecological measures used for each species.

Ecological measure Definition
Environmental volume   Three-dimensional range of a species in an ordination space based on 53 environmental measures for each of 92 plots.
 The range on each axis was the distance of the sites with the largest and smallest axis scores that contained that species.
Abundance   Mean cover value of a species, based on a six-point Braun-Blanquet scale, for all plots in which the species appeared.
Ecological breadth   Classification of each species into one of four groups based on its general distribution: wet-grassland specialist (WGS),
 wet-grassland generalist (WGG), mesic-habitat generalist (MHG), and generalist (G).
  Wet-grassland specialists   Species characteristic of Iberian Atlantic climates.
  Wet-grassland generalists   Species that are found in a range of mesic habitats.
  Mesic-habitat generalists    Species that are found in a range of mesic habitats.
  Generalists    Species that are typically found in nonwetland habitats.
Habitat occupancy    The percentage of plots within an environmental volume that were occupied by a given species.
Regional range    The extent of a species' range within the Iberian Peninsula, in km2.
Continental range    The number of geographic units across Europe and the Mediterranean basin with a recorded presence for a species.
 


Habitat specificity was measured at two scales: environmental volume and ecological breadth. Environmental volume measured habitat specificity within the set of sample plots in the wet grasslands. Ecological breadth measured it across a species' range. Environmental volume was calculated as follows. In each plot, we recorded 53 variables (Appendix 3) that were ultimately reduced into a three-dimensional, multivariate space, the first three eigenvectors from a principal components analysis. For each plant species, the environmental volume was the rectangular solid determined by the maximum and minimum coordinates on all three axes. See Appendix 4 for details.

For ecological breadth, each species was classified into one of four groups: wet-grassland specialist (WGS), wet-grassland generalist (WGG), mesic-habitat generalist (MHG), and generalist (G). The first three groups consist of species whose typical habitat is contained within the group of sampled plots. Wet-grassland specialists are characteristic of Atlantic climates. In our study, they were found only in the high-elevation plots because those plots receive the highest amounts of precipitation and have the lowest potential evapotranspiration. Wet-grassland generalists are potentially found in all of the sampled sites, but would not be found in other habitats. Mesic-habitat generalists are also found in other mesic habitats such as deciduous forests (Quercus pyrenaica associations) and coniferous forests (Pinus sylvestris associations). Generalist species are typically not found in these habitats and, thus, are at the edge of their habitat ranges. This classification scheme was made independently of the data and was based on the flora and habitat descriptions in Rivas Martínez (1963), Tutin et al. (1964, 1968, 1972, 1976, 1980), Bolós and Vigo (1984, 1990, 1996), Valdés et al. (1987), and Fernández González (1991).

Habitat occupancy was based on each species' environmental volume. For each species, the number of plots within its volume was determined. Habitat occupancy was calculated as the percentage of plots within the volume that were occupied. This measure assumes that, for each species, all habitats intermediate to those already occupied are suitable.

Geographic range was measured at two scales: regionally across the Iberian Peninsula, and continentally across Europe and the Mediterranean basin. For both measures, we used published distribution maps and lists. At the regional scale, geographic range was measured as the extent of the range within the Iberian Peninsula in km2. These regional ranges were taken from the maps of Bolós and Vigo (1984, 1990, 1996) and Hultén and Fries (1986) and were determined by scanning the maps using a Tamaya Digitizing Area-Line Meter (Tamaya Technics, Tokyo, Japan). At the continental scale, geographic range was measured as the number of geographic units with a recorded presence for the species. Data were from Tutin et al. (1964, 1968, 1972, 1976, 1980), Hultén and Fries (1986), and Greuter et al. (1984). These sources define 49 geographic units that are variously entire countries, groups of countries, or smaller units such as discrete islands. Although this measure of continental range does not correct for size differences among geographic units, it suffices as a general measure of continental range.

Data were analyzed using Systat (SYSTAT 6.1 for Windows, SPSS, Chicago, Illinois, USA). The measures of local abundance, environmental volume, and habitat occupancy were not normally distributed. Thus, all analyses were done as nonparametric tests.


RESULTS

Local characteristics

The distribution of site occupancy by species shows a typical log-normal distribution, with 29% of the species in one plot only and 17% of the species in two plots. Most species had low local abundances and small environmental volumes (Fig. 3). On average, species occupied about half of the plots within their environmental volumes. We next examine the relationships among the variables and refer the reader to Table 2 for predictions of the two theories.


Fig. 3. Frequency histograms of (A) mean abundance, (B) environmental volume, (C) habitat occupancy, (D) regional range, and (E) continental range for 237 plant species sampled in the wet grasslands of the Sierra de Guadarrama. In (A) and (B), solid bars correspond to species with 100% of their environmental volumes occupied; open bars correspond to species with <100% of their environmental volumes occupied.

GIF image file (7 K)


Local abundance and environmental volume were positively correlated (Fig. 4A). The situation was more complicated for correlations with habitat occupancy, however. When all species were included in the analyses, both local abundance and environmental volume were negatively correlated with habitat occupancy (Figs. 4B,C). The negative correlations were due, however, to the large number of species found in only one or two sites. A species that appears in a single plot perforce must occupy 100% of its volume. Similarly, most species found in only two plots, and occasionally in three plots, also had 100% occupancies. This large group of species had the potential to distort the analysis. When those 94 species were removed, the correlation with habitat occupancy became positive for local abundance (rS = 0.20, P < 0.02, n = 138) and disappeared for environmental volume (rS = 0.02, P > 0.5, n = 138). The species found in only one or two plots had a range of abundances from small to large. Otherwise, species with broader local habitat specificities tended to have higher abundances where present.


Fig. 4. Relationships among mean abundance, environmental volume, and habitat occupancy. Symbols indicate the ecological breadth group: triangle, WGS; circle, WGG; square, MHG; and diamond, G. Solid symbols correspond to those species with 100% of their environmental volumes occupied; open symbols correspond to those species with < 100% of their environmental volumes occupied. Correlations and statistical significance are given in Table 5.

GIF image file (6 K)


Ecological breadth groups

The relationship between these three traits (environmental volume, local abundance, and habitat occupancy) and ecological breadth was more complex (Table 4). Again, see Table 2 for the theory predictions. First, we considered environmental volume, which tells us whether we were successful in our classification of specialists and generalists. As expected, on average, mesic- habitat generalists had the largest volumes, followed by wet-grassland generalists, then generalists, and finally wet-grassland specialists.


Table 4. Species characteristics of environmental volume, local abundance, habitat occupancy, and geographic range of Spanish montane grassland species as a function of their ecological breadth (WGS, wet-grassland specialists; WGG, wet-grassland generalists; MHG, mesic-habitat generalists; G, generalists; defined in Table 3). Volumes are expressed as a percentage of the maximum possible. Differences among groups were determined by a Kruskal-Wallis nonparametric ANOVA and Dunn's nonparametric post hoc test (Zar 1984: 200). Statistically significant (alpha = 0.05) differences among groups are indicated by different superscripts.

 
Ecological breadth group  Statistics 
WGS  WGG  MHG  Chi2  P 
Environmental volume 
median 0.00b  2.76a  6.65a  0.48ab   21.75   0.0001 
range (0.0-21)  (0.0-75)  (0.0-75)  (0.0-63) 
no. species 28  71  66  67 
Mean abundance 
median 1.00a  1.10a  1.00a  0.50b  39.70  0.0001 
range (0.1-2.0)  (0.1-4.0)  (0.1-2.5)  (0.1-2.0) 
no. species 28  71  66  67 
Habitat occupancy 
median 100.0  55.2  55.4  57.1  6.84  0.08 
range (18-100)  (14-100)  (11-100)  (15-100) 
no. species 28  71  66  67 
Regional range (x105 km2
median 1.40c  2.19bc  2.80ab  3.81a  18.02  0.0001 
range (0.08-5.81)  (0.25-5.81)  (0.29-5.81)  (0.02-5.81) 
no. species 28  66  53  52 
Continental range 
median 33  33  34  32  4.16  0.24 
range (7-48)  (3-48)  (4-47)  (2-48) 
no. species 27  56  48  40 


For mean abundance, generalists had significantly lower mean abundances than species in the other three categories. Of the generalists, 36% had the lowest mean abundance possible, 0.1. Thus, these habitats were marginal for the generalist species, as might be expected.

Habitat occupancy did not differ among groups. This trait was unrelated to habitat specificity. When species with 100% habitat occupancy were eliminated, median occupancy was 27%, 44%, 48%, and 40% for the four groups, respectively. Again, they did not differ (chi-square = 4.83, P = 0.18).

Next, we examined correlations among local abundance, environmental volume, and habitat occupancy within each group. When all species were included, all correlations had the same sign as the overall correlations (Table 5). Local abundance and environmental volume were positively correlated, whereas both were negatively correlated with habitat occupancy. The only notable variation among groups was the substantially lower correlation of local abundance and environmental volume for wet-grassland specialists. Thus, in general, the ecological specificity of a species did not alter the tendency for broader local habitat specificity to be associated with higher abundance.


Table 5. Spearman rank correlations among local abundance, environmental volume, habitat occupancy, and regional ranges across over all species (n = 232) and for groups of Spanish montane grassland species based on their ecological breadth, as defined in Table 3. For the overall correlations, "Excluded" indicates species with 100% occupancy deleted (n = 138). Groups are as in Table 3. Correlations that differ significantly from zero (alpha = 0.05) are indicated in boldface. Sample sizes for ecological breadth groups are given in Table 2.


Overall Ecological breadth group
Traits

All

Excluded

WGS

WGG

MHG

G

Abundance-volume

0.41

0.33

0.02

0.40

0.47

0.57

Abundance-occupancy

-0.22

0.20

-0.10

-0.17

-0.18

-0.46

Abundance-regional range

-0.09

-0.12

-0.05

-0.07

0.12

0.16

Abundance-continental range

0.17

0.07

-0.07

0.11

0.15

0.44

Volume-occupancy

-0.75

0.02

-0.86

-0.69

-0.58

-0.83

Volume-regional range

0.24

0.14

0.13

0.05

0.41

0.27

Volume-continental range

0.28

0.14

0.08

-0.09

0.42

0.51

Occupancy-regional range

-0.26

-0.05

-0.40

-0.09

-0.29

-0.29

Occupancy-continental range

-0.24

-0.04

-0.06

0.01

-0.34

-0.42

Regional-continental range

0.46

0.64

0.33

0.35

0.64

0.54


Geographic range

We next considered the two measures of geographic range: regional and continental. The two measures were correlated with each other (rS = 0.46, P < 0.001, n = 163; Fig. 5). Both regional and continental range were positively correlated with environmental volume and negatively correlated with habitat occupancy (Figs. 6 and 7). In contrast, for local abundance, regional range was not correlated, whereas continental range was positively correlated. All of these correlations were nonsignificant when the species with 100% occupancy were removed, although the signs of the correlations remained the same. For the other measure of habitat specificity, ecological breadth, the rank order with regard to regional range was as expected. Specialist species had the smallest regional ranges, followed by wet-grassland generalists, then mesic-habitat generalists, and, finally, generalist species had the largest ranges (Table 4). The groups did not differ, however, in continental ranges.


Fig. 5. Relationship between regional range and continental range. Correlations and statistical significance are given in Table 5.

GIF image file (4 K)


Fig. 6. Relationships between regional range and (A) mean abundance, (B) environmental volume, and (C) habitat occupancy. Correlations and statistical significance are given in Table 5.

GIF image file (8 K)


Fig. 7. Relationships between continental range and (A) mean abundance, (B) environmental volume, and (C) habitat occupancy. Correlations and statistical significance are given in Table 5.

GIF image file (8 K)


Categorization

To explore more completely the patterns of commonness and rarity, we dichotomized the data to fit within the scheme shown in Table 1B. Although such a procedure subsumes information within the categories, it allows a multivariate view of the data. We dichotomized the data using the approximate median of each distribution: geographic range wide = present in at least 35 geographic units, habitat specificity broad = environmental volume at least 11% of total, habitat occupancy high = more than 40%, local abundance large = maximal abundance at least 2.

Species were not evenly distributed among the 16 categories (Table 6), being overrepresented among categories that included high habitat occupancy. The most frequent category was endemic indicator species with small abundances (36%), followed by common species. The least frequent categories were the four with a broad habitat specificity and small local abundance. Two additional categories differing in local abundance (wide geographic range, restricted habitat specificity, and low habitat occupancy) also included few species. Only one category (narrow geographic range, broad habitat specificity, small abundance, and high occupancy) had no species.


Table 6. Percentage of species sampled in montane wet grasslands that were found within each category of our scheme described in Table 1B. The upper value is for all species (n = 220) and the lower value is for wet-grassland specialists and generalists only (n = 96). See text for the threshold values that were used to dichotomize the categories.



 
Geographic range  Wide  Narrow 
Habitat specificity  Broad  Restricted  Broad  Restricted 
Abundance  Large  Small  Large  Small  Large  Small  Large  Small 
Habitat occupancy high  9.5  0.9  5.0  7.7  7.3  0.0  8.2  35.5 
9.4  1.0  7.3  11.5  8.3  0.0  12.5  28.1 
Habitat occupancy low  4.5  1.4  2.2  2.7  5.0  1.8  4.5  3.6 
4.2  0.0  1.0  1.0  4.2  1.0  7.3  3.1 
 


There might be a problem, however, with using all of the species for this analysis. Species with habitat preferences outside of wet meadows (e.g., trees, xerophytes), would appear in this categorization as habitat specialists. Thus, we repeated the previous analysis using only those species classified as wet-grassland specialists or generalists. The overall pattern was similar, with a few exceptions. First, the endemic indicator species were more likely to have large than small abundances, although the total frequency of this category remained about the same. Second, there were increases in the frequency of species classified both as endangered and as indicators.


DISCUSSION

In general, there was a tendency for all species characteristics to be positively correlated. In other words, all possible combinations of traits shown in Table 1B were not found or were infrequent. Species tended either to have wide geographic ranges, broad habitat distributions, large local abundances, and high habitat occupancies or to have the opposite combination of traits. However, these were only general trends, and overall correlations were generally low to intermediate. For example, there were several species with restricted habitat distributions and large local abundances.These results suggest the existence of both superior species and specialists. A good example of a specialist is Lysimachia vulgaris. This species was found in just one plot, but in that plot it was by far the most abundant species.

As in a previous study of the British flora (Rabinowitz et al. 1986), we found little evidence for species with both broad habitat specificities and small abundances. The categories with the fewest species had either small mean abundances or low habitat occupancies. Such species will have the greatest vulnerability to demographic stochasticity (Lande 1988). Thus, such species may be infrequent because they tend to go extinct more often, rather than because they cannot evolve.

Conservation implications

Among the sampled species, none is officially listed as endangered in Iberia (Gómez-Campo 1987). However, three species (Eleocharis quinqueflora, Ilex aquifolium, and Juncus bulbosus) are included in the official catalog of endangered species of the Madrid regional government (CAM 1992). Our analysis classified the first two species as endemic indicators. Of the 28 plants within our endangered categories, Cardamine pratensis, Carex panicea, and Juncus bulbosus are actually rare species that could be considered as endangered in the studied region. Only one of them, however, is now officially considered vulnerable. Other species originally classified as endangered in our scheme were rare in wet grasslands only because they are woody species or species of other habitats such as xerophytes or mesophytes. Thus, an analysis such as ours is a useful first step in conservation efforts.

Tests of the theories

Support for the two theories was mixed, with evidence supporting and refuting both. Consider the predictions laid out in Table 2. As predicted by the trade-off theory, the most frequent category was "endemic indicators." However, unlike the prediction for this theory, the "sparse" category had few species. As predicted by the superior organism category, the next most frequent categories were "common" and "endangered."

Geographic range, habitat specificity, and local abundance were all positively correlated in agreement with the superior organism theory, but not with the trade-off theory. On the other hand, the trade-off theory predicts the existence of species with very small volumes, high local abundance, and high occupancies. Such species were found in our samples (e.g., Lysimachia vulgaris and Scirpus lacustris).

With respect to the ecological groups, the predictions of neither theory were upheld with respect to local abundance and habitat occupancy (Tables 2 and 4). For environmental volume, the ordering of groups was consistent with the trade-off theory, although the differences were not statistically significant. Only for geographic range, based on distributions within Iberia, were the predictions upheld. Unfortunately, both theories make the same prediction, so these results fail to discriminate between them.

How general are our conclusions? Nearly all of the species in these plots were herbaceous. Trees and shrubs were nearly absent and always strays. Woody plants may differ in their patterns of commonness and rarity. In addition, we sampled only one habitat type, montane wet grasslands. On the other hand, patterns were generally strong across a large number of species and measures. For example, results concerning geographic ranges were consistent at both the regional and continental scales.

Evidence for the superior organism theory from other studies is mixed. A study of plant species distributions (Scheiner and Rey Benayas 1997) found a positive correlation between local abundance and habitat occupancy in 15 of 15 landscapes from across the globe. On the other hand, several reviews of animal species distributions (Gaston 1994, 1997, Lawton et al. 1994) found a positive correlation between geographic range and local abundance in only some studies, albeit the majority of them.

Speculations

An open question is how both superior species and habitat specialists can coexist. Superior species should outcompete specialists because of their greater habitat breadth. Why has this not occurred? It may be that the local populations of the superior species are not self-sustaining. They may be sink populations. This explanation is bolstered by the observation that the generalist species, which have the largest regional ranges, also have the lowest average local abundances (Table 4). These species may have superior dispersal abilities, creating the appearance of local dominance, whereas the specialists species are still able to outcompete them within their local niches. The mechanisms governing that competition are varied, including responses to herbivory, resource fluctuation, and disturbance. Further, detailed studies are necessary to discover their workings.

We recognize that this explanation belongs to that class of ecological/evolutionary explanations of the sort "there has to be a trade-off somewhere." We (speaking collectively of ecologists and evolutionary biologists) regularly retreat to such explanations because we are otherwise unable to account for the persistence of diversity in the world. We recognize, though, that a "well it has to be there somewhere" defense is ultimately unsatisfactory. The resolution of this dilemma may lie in a close study of the apparently superior species to discover if and how we are being mislead as to their nature. Three of those species (Plantago lanceolata, P. major, and Trifolium repens) are widespread weeds in North America. As such, they tend to be found in disturbed habitats, which is consistent with our "sink population" explanation.

Missing from either theory is an explanation of why a particular species is common or rare. Two recent reviews (Gaston 1994, Kunin 1997) list a number of causes of rarity. The proximal mechanisms reduce to three categories: ecological specialization, lack of dispersal, and historical contingency. Our data suggest that ecological specialization may be an important cause of rarity. Species with small environmental volumes tend to have lower abundances and, particularly, smaller geographic ranges. Whether this specialization is due to trade-offs or to being an inferior species is an open question. We speculate, however, that the answer is neither one nor the other. Both mechanisms probably play a role.

Further dissection of these categories is instructive (e.g., Rosenzweig and Lomolino 1997, and other chapters in Kunin and Gaston 1997) because they examine nonproximal mechanisms, those that cause ecological specialization. Among the suggested mechanisms are: breeding systems that favor selfing, low reproductive investment, low amounts of genetic variation, low population densities, and chaotic population dynamics. For animals, some of these syndromes are associated with larger body sizes (Gaston 1994).

In another analysis, Kelly (Kelly 1996, Kelly and Woodward 1996) searched for ecological correlates of commonness and rarity in the floras of Great Britain and Crete. She asked whether there are particular attributes of species that give them wide geographic ranges. Geographic range was defined similarly to our regional measure, the number of counties or 10 km2 blocks in which a species appeared. She looked at a number of attributes, including life-form, pollination syndrome, and dispersion mechanism. Only life-form (trees were more common than shrubs) and pollination syndrome (wind-pollinated plants were more common than animal-pollinated plants) were found to explain geographic range. Unclear is the generality of these results. Both regions are islands. Will continental floras differ?

Our data do not address the dispersal issue. However, this mechanism implies that habitats contain less than the maximal number of species that they could hold. That is, if a species is rare only because it is unable to get to more suitable habitats, then other mechanisms such as competitive exclusion are not at work. Competitive exclusion would fall under the heading of rarity due to ecological specialization. Is there evidence for local areas with fewer species than might be expected based on factors such as levels of productivity? Yes. Two studies (Pärtel et al. 1996, Caley and Schluter 1997) found that local diversity was correlated with regional diversity. They (and we) interpret these results to suggest that local diversity is limited, in part, by the size of the regional species pool.

Thus, patterns of commonness and rarity appear to be linked with patterns of high and low diversity. These two aspects of species distributions are complementary and represent the patterns of column totals and row totals in a site-species matrix (e.g., Appendix 1). Typically, these two aspects of species distributions are studied separately. We suggest that integrating such studies will provide additional insights into both issues.


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Acknowledgments:

We thank several anonymous reviewers for comments on a previous draft of this manuscript. Funding was provided by the projects "Humedales en áreas de descarga de acuíferos en territorios graníticos (Sierra de Guadarrama)" (Comunidad Autónoma de Madrid nº C129/91), and "Determinantes de la diversidad biológica en ecosistemas mediterraneós. Papel de los procesos locales y regionales" (DGICYT). Additional funding was provided by the Universidad de Alcalá for travel by S. M. Scheiner.


LITERATURE CITED

Bernáldez, F. G., J. M. Rey Benayas, and A. Martínez. 1993. Ecological impact typology on wetlands produced by groundwater extraction (Douro River, Spain). Journal of Hydrology 141:219-238.

Bolós, O., and J. Vigo. 1984, 1990, 1996. Flora dels Paisös Catalans. Vols. I-II-III. Edition Barcino, Barcelona, Spain.

Brown, J. H. 1984. On the relationship between abundance and distribution of species. American Naturalist 124:255-279.

Brown, J. H., D. W. Mehlman, and G. C. Stevens. 1995. Spatial variation in abundance. Ecology 76:2028-2043.

Burgman, M. A. 1989. The habitat volumes of scarce and ubiquitous plants: a test of the model of environmental control. American Naturalist 133:228-239.

Caley, M. J., and D. Schluter. 1997. The relationship between local and regional diversity. Ecology 78:70-80.

CAM (Comunidad Autónoma de Madrid). 1992. Catálogo regional de especies amenzadas de fauna y flora silvestres. Boletín Oficial de la Comunidad Autónoma de Madrid, Numero186; 82 y 85, Madrid, Spain.

Fernández González, F. 1991. La vegetación del Valle del Paular (Sierra de Guadarrama, Madrid). Lazaroa 12:153-272.

Fox, L. R., and P. A. Morrow. 1981. Specialization: species property or local phenomenon? Science 211:887-893.

Futuyuma, D. J., and G. Moreno. 1988. The evolution of ecological specialization. Annual Review of Ecology and Systematics 19:207-233.

García Sánchez-Colomer, M. 1998. Variación del ambiente y su efecto en las comunidades de plantas de los humedales de la Sierra de Guadarrama. Dissertation. Universidad Autónoma de Madrid, Spain.

Gaston, K. J. 1994. Rarity. Chapman and Hall, London, UK.

_______ . 1997. What is rarity? Pages 30-47 in W. E. Kunin and K. J. Gaston, editors. The biology of rarity. Causes and consequences of rare-common differences. Chapman and Hall, London, UK.

Gaston, K. J., and J. H. Lawton. 1990. Effects of scale and habitat on the relationship between regional distribution and local abundance. Oikos 58:329-335.

Gómez-Campo, C., editor. 1987. Libro rojo de especies vegetales amenazadas de España peninsular e Islas Baleares. ICONA, Madrid, Spain.

Greuter, W., H. M. Burdet, and G. Long, editors. 1984. MED-CHECKLIST. A critical inventory of vascular plants of the circum-mediterranean countries. Volumes 1, 2, and 3. Editions des Conservatoire et Jardin botaniques de la Ville de Genéve, Switzerland.

Hughes, L., E. M. Cawsey, and M. Westoby. 1996. Geographic and climatic range sizes of Australian eucalyptus and a test of Rapoport's rule. Global Ecology and Biogeography Letters 5:128-142.

Hultén, E., and M. Fries. 1986. Atlas of North European vascular plants north of the Tropic of Cancer. 3 volumes. Koeltz Scientific Books, Königsstein, Germany.

Hutchinson, G. E. 1957. Concluding remarks. Cold Spring Harbor Symposium on Quantitative Biology 22:415-427.

Kelly, C. K. 1996. Identifying plant functional types using floristic databases: ecological correlates of plant range size. Journal of Vegetation Science 7:417-424.

Kelly, C. K., and F. I. Woodward. 1996. Ecological correlates of plant range size: taxonomies and phylogenies in the study of plant commonness and rarity in Great Britain. Philosophical Transactions of the Royal Society of LondonB 351:1261-1269.

Kunin, W. E. 1997. Introduction: on the causes and consequences of rare-common differences. Pages 3-11 in W. E. Kunin and K. J. Gaston, editors. The biology of rarity. Causes and consequences of rare-common differences. Chapman and Hall, London, UK.

Kunin, W. E., and K. J. Gaston. 1997. The biology of rarity. Causes and consequences of rare-common differences. Chapman and Hall, London, UK.

Lande, R. 1988. Genetics and demography in biological conservation. Science 241:145-146.

Lawton, J. H., S. Nee, A. J. Letcher, and P. H. Harvey. 1994. Animal distributions: patterns and processes. Pages 41-58 in P. J. Edwards, R. M. May, and N. R. Webb, editors. Large-scale ecology and conservation biology. Blackwell Scientific, Oxford, UK.

McCune, B., and M. J. Mefford. 1995. PC-ORD. Multivariate analysis of ecological data. Version 2.0. MjM Software Design, Gleneden Beach, Oregon, USA.

McGuiness, K. A. 1984. Equations and explanations in the study of species-area curves. Biological Review 59:423-440.

Myers, A. A., and P. S. Giller, editors. 1988. Analytical biogeography. An integrated approach to the study of animal and plant distribution. Chapman and Hall, London, UK.

Orians, G. H. 1997. Evolved consequences of rarity. Pages 190-208 in W. E. Kunin and K. J. Gaston, editors. The biology of rarity. Causes and consequences of rare-common differences. Chapman and Hall, London, UK.

Pagel, M. D., R. M. May, and A. Collie. 1991. Ecological aspects of the geographic distribution and diversity of mammal species. American Naturalist 137:791-815.

Pärtel, M., M. Zobel, K. Zobel, and E. van der Maarel. 1996. The species pool and its relation to species richness: evidence from Estonian plant communities. Oikos 75:111-117.

Rabinowitz, D. 1981. Seven forms of rarity. Pages 205-217 in H. Synge, editor. The biological aspects of rare plant conservation. John Wiley, Chichester, UK.

Rabinowitz, D., S. Cairns, and T. Dillon. 1986. Seven forms of rarity and their frequency in the flora of the British Isles. Pages 182-204 in M. E. Soulé, editor. Conservation biology: the science of scarcity and diversity. Sinauer, Sunderland, Massachusetts, USA.

Rapoport, E. H. 1982. Areography: geographical strategies of species. Pergamon, Oxford, UK.

Rey Benayas, J. M., M. G. S. Colomer, C. Levassor, and I. Vázquez-Dodero. 1998. The role of wet grasslands in biological conservation in Mediterranean landscapes. Pages 61-72 in C. Joyce and M. Wade, editors. John Wiley, New York, New York, USA.

Rivas Martínez, S. 1963. Estudio de la vegetación y flora de las sierras de Guadarrama y Gredos. Anales del Jardín Botánico de Madrid 21-I:1-325.

Rosenzweig, M. L., and M. V. Lomolino. 1997. Who gets the short bits of the broken stick? Pages 63-90 in W. E. Kunin and K. J. Gaston, editors. The biology of rarity. Causes and consequences of rare-common differences. Chapman and Hall, London, UK.

Scheiner, S. M., and J. M. Rey Benayas. 1997. Placing empirical limits on metapopulation models for terrestrial plants. Evolutionary Ecology 11:275-288.

Stevens, G. C. 1989. The latitudinal gradient in geographic range: how so many species coexist in the tropics. American Naturalist 133:240-246.

Tutin, T. G., V. H. Heywood, N. A. Burges, D. M. Moore, D. H. Valentine, S. M. Walters, and D. A. Webb, editors. 1964. Flora Europaea. Volume I. Cambridge University Press, Cambridge, UK.

Tutin, T. G., V. H. Heywood, N. A. Burges, D. M. Moore, D. H. Valentine, S. M. Walters, and D. A. Webb, editors. 1968. Flora Europaea. Volume II. Cambridge University Press, Cambridge, UK.

Tutin, T. G., V. H. Heywood, N. A. Burges, D. M. Moore, D. H. Valentine, S. M. Walters, and D. A. Webb, editors. 1972. Flora Europaea. Volume III. Cambridge University Press, Cambridge, UK.

Tutin, T. G., V. H. Heywood, N. A. Burges, D. M. Moore, D. H. Valentine, S. M. Walters, and D. A. Webb, editors. 1976. Flora Europaea. Volume IV. Cambridge University Press, Cambridge, UK.

Tutin, T. G., V. H. Heywood, N. A. Burges, D. M. Moore, D. H. Valentine, S. M. Walters, and D. A. Webb, editors. 1980. Flora Europaea. Volume V. Cambridge University Press, Cambridge, UK.

Valdés, B., S. Talavera, and E. Fernández Galiano. 1987. Flora vascular de Andalucía Occidental. 3 Volumes. Ketres Edition, Barcelona, Spain.


APPENDICES

Appendix 1. Site-by-species data matrix showing the abundances of the 232 species in the 92 plots. Nomenclature follows Tutin et al. (1964, 1968, 1972, 1976, 1980).

A tab-delimited version of this appendix is available as a text file.

                                                                                                                                                                                                                                                                                                                                                                                       PLOT
        PLANTS                                          1       2       3       4       5       6       7       8       9       10      11      12      13      14      15      16      17      18      19      20      21      22      23      24      25      26      27      28      29      30      31      32      33      34      35      36      37      38      39      40      41      42      43      44      45      46      47      48      49      50      51      52      53      54      55      56      57      58      59      60      61      62      63      64      65      66      67      68      69      70      71      72      73      74      75      76      77      78      79      80      81      82      83      84      85      86      87      88      89      90      91      92
1       Achillea millefolium L.                         0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
2       Adenocarpus hispanicus (Lam.) DC.               0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0
3       Agrimonia eupatoria L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0
4       Agrostis canina L.                              0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       2       1       0       2       1       0       0       0       0.1     0       0       0       0       1       0       1       0.1     0       2       2       0       2       1       2       0       4       1       0       0       0       0       0       0       0       0       2       1       2       0       1       0       1       0       0       0       0       0       2       3       0       0       0       0       0       0       0       0       0       3       0       3       1       0       0       0       0       0       0       0       0       0       1       1       2       1       1       0
5       Agrostis capillaris L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0
6       Agrostis castellana Boiss. & Reuter             2       4       4       4       3       4       5       3       4       3       4       2       1       1       0       1       1       1       1       1       1       1       0.1     0.1     0.1     4       3       3       0       0.1     0.1     5       4       1       0       1       0       1       1       0       0.1     2       0       4       1       4       4       5       5       2       0       0       0       0       0       2       2       1       0       2       1       1       0       0       2       0.1     2       0       0       0       0       0       4       0       2       0       0       0       3       1       1       1       3       0       1       1       0       0       0       0       0       0
7       Agrostis stolonifera L.                         0       0       0       0       0       0       0       0       0       0.1     1       2       1       1       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0.1     0       0       0       0       0       1       0       0       0       0       0       0
8       Alchemilla filicaulis Buser                     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
9       Alliaria petiolata (Bieb.) Cav. & Grand.        0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
10      Allium oleraceum L.                             0       0       0       0       0       0.1     0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
11      Allium sphaerocephalon L.                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
12      Alnus glutinosa (L.) Gaertner                   0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
13      Alopecurus arundinaceus Poiret                  0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       1       0       0       0       0       1       0       0       1       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0.1     2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0
14      Alopecurus geniculatus L.                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0.1     0       0       0       0       0       0
15      Anagallis tenella (L.) L.                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       1       1       0       0       1       3       0       0       2       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
16      Anarrhinum bellidifolium (L.) Willd.            0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
17      Anthoxanthum odoratum L.                        0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       1       1       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       1       0.1     1       0       0       1       0       1       1       0       0       0       0       0       0       0       1       0       1       1       1       2       1       1       1       1       0       1       1       0.1     1       0       0       0       0       0       0       0       0       2       2       1       2       1       1       0       0       0       0       1       0       0       0       0       1       0       2       0       1
18      Apium nodiflorum (L.) Lag.*                     0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
19      Aristolochia fontanesii Boiss. & Reut.          0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
20      Arrhenatherum album (Vahl) W.D. Clayton         0       0       0       0.1     1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       1       0       0       0.1     0       0       0       0       0       0       0       0       0       0
21      Arrhenatherum elatius (L.) Beauv. ex C. Presl.  0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0
22      Asphodelus ramosus L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       1       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
23      Baldellia ranunculoides (L.) Parl.              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
24      Bellis perennis L.                              0       4       2       0       0.1     0       0.1     0.1     0       1       1       2       0       0       2       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       2       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
25      Betula pubescens Ehrh.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       4       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       1       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
26      Brachypodium sylvaticum (Hudson) Beauv.         0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       1       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0
27      Briza media L.                                  0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0.1     0       0       1       0       0       0       0       1       0       1       0       1       0       0       2       1       1       0       0       0       0.1     0.1     0       0       1       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       1       1       0       0.1     0       0       1       0       0       0       0       0       0       0       0       1       0       0       0.1     0       1       0       0       0       0       1       0       0       0       0       0       0       0       0       0
28      Bryonia cretica L.                              0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
29      Calluna vulgaris (L.) Hull.                     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0
30      Campanula rapunculus L.                         0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
31      Cardamine pratensis L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
32      Carduncellus araneosus Boiss. & Reuter          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
33      Carduus carpetanus Boiss. & Reuter              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0
34      Carex binervis Sm.                              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0.1     0       1       1       0       0       0.1     0.1     2       0       1       2       2       2       3       2       0       2       1       3       0       1       0       1       4       1       0       0       0       0       0       1       0       0       0       0       0       1       0.1     0.1     1       2       2       0       1       0       0       0.1     0       0       0       0       0       0       0       0       1       0       1       0       1       2       0       0       2       3       1       0       0       0       0       0       0       0       0       0
35      Carex demissa Hornem.                           0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0.1     0       0       0       0       0       0       1       1       0       1       0       3       3       1       1       0       2       1       0       2       0       2       0       0       1       0       0       0       0       0       0       0       0       1       0       1       2       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       2       0       0       0       0       0       0       0       0       0       0       0.1     2       2       2       0
36      Carex divisa Hudson                             0       3       2       2       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       2       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       2       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       1       0       0       1       0       0       0       0       0       0       0       0       0       0
37      Carex divulsa Stokes                            0       0       0       1       1       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
38      Carex echinata Murray                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0.1     0       1       0       1       0       0       1       0       0       0       0       0       0       0       2       2       1       1       3       1       1       0       0       3       0       0       0.1     2       0       0       0       0       0       0       0       0       0       0       0       1       2       1       0       0       0       0       0       0       0       0       0       4       3       2       1       3       1
39      Carex flacca Schreb.                            0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       1       0.1     0       0       1       0       0       0.1     1       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0
40      Carex hirta L.                                  0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       1       0       0       0       0       0       0
41      Carex muricata L. subsp. lamprocarpa Celak      0       0       0       1       0.1     0.1     0.1     0.1     0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0.1     0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0
42      Carex nigra (L.) Reich.                         0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       1       0       0       0       1       1       0       1       0       0       1       0       0       0       0       0       0       0       0       0       0       1       0       3       2       4       2       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       1       0       0       1       0       0       0       0       0       0       0       0       0       0       4       4       3       3       4       2
43      Carex ovalis Good.                              0       0       0       0       0       0.1     0       0       0       1       1       1       2       1       0       0       0       0       3       2       0       0       1       2       0       0       1       2       0       0       1       0       0       0       1       1       1       0       0       0.1     0       0       0       0       2       0       0       0       0       0       2       0       0       1       0       1       1       2       1       1       1       1       0       0       0       2       1       0       0       0       0       0       0       3       1       0       0       1       1       1       1       0       0       0       1       2       0       0       0       0       0       0
44      Carex panicea L.                                0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       2       2       2       0       0       2       0       0       0       0       0       0       0       1       0       0       0       0       0.1     0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       3       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0
45      Carlina corymbosa L.                            0       0.1     0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
46      Carlina racemosa L.                             0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
47      Carum verticillatum (L.) Koch                   0       0       0       0       0       0       0       0       0       0       2       0       1       1       0.1     0.1     0.1     0       2       1       0.1     1       1       0       1       0       0.1     1       3       2       0       1       0       3       3       2       2       3       1       0       2       1       0.1     0       0       0       0       0       0       3       0       0.1     1       1       0.1     1       2       2       0       2       2       0       1       0       1       3       2       0       4       5       0.1     0       0       0       3       3       0       0       0       1       4       2       0       0       0       2       0       0       0       1       0       4
48      Centaurea collina L.                            0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
49      Centaurea nigra L.                              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       1       0       0       0       0       0       0       0       0       0
50      Centaurea scabiosa L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
51      Cerastium fontanum Baumg.                       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0.1     1       0       0       0       0       0       1       0.1     0       0       0       0       0       0       0       0       0.1     0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0.1     0.1     0       0.1     0.1     0.1     0.1     1       0       0       0       1       0       0.1     0       0       0       0       0       0       0       0       1       1       1       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
52      Chaerophyllum hirsutum L.                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
53      Chamaemelum nobile (L.) All.                    0       2       2       0       0       0       0       0       0       2       2       1       0.1     0       0       0       0       1       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
54      Cirsium flavispina Boiss. ex DC.                0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
55      Cirsium palustre (L.) Scop.                     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
56      Clinopodium vulgare L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0
57      Convolvulus arvensis L.                         1       0       1       1       0       3       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       4       0.1     0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0
58      Corrigiola litoralis L.                         0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
59      Crataegus monogyna Jacq.                        0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0.1     0       0       0       0       0       0
60      Crepis vesicaria L.                             0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
61      Cruciata glabra (L.) Ehr.                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
62      Cucubalus baccifer L.                           0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
63      Cynodon dactylon (L.) Pers.                     0       4       3       4       0       1       0       0       1       0       0       0       0       0       0       0       0       0       0       0       4       4       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       4       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       1       2       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
64      Cynosurus cristatus L.                          0       0.1     0.1     0       0       2       0       2       0       1       1       1       2       1       1       2       2       0       0       1       3       4       1       1       1       0       1       2       2       1       2       1       0       2       0       3       2       3       0       0       3       0       0       0       0       1       0       1       2       0       2       0       0       0       0       3       1       3       4       3       0       0.1     0       3       2       2       1       0       0       0       0       4       2       2       2       0       1       0       1       1       2       2       1       0       0.1     1       0       0       0       0       0       0
65      Cyperus longus L.                               0       1       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       1       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0.1     0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
66      Cytisus purgans (L.) Boiss.                     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
67      Dactylis glomerata L. subsp. (Roth) Nyman       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
68      Dactylorhiza elata(Poiter)Soó ssp.sesquipedalis(Willd.)Soó       1       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       1       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0
69      Dactylorhiza sambucina (L.) Soó                 0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0.1     0       0       0.1     0       0       0       0       0       0       0       0       0       0       0
70      Danthonia decumbens (L.) DC.                    0       0       0       0       0       0       0       0       0       1       1       1       2       1       0       2       1       0       0       1       0       0       0       0       1       0       1       1       0.1     0       0.1     1       1       3       0       1       0.1     0       0       0       0       1       0       0       0       0       0       1       0       0.1     0       0       0       0       0       0       0       0       0       1       1       0       0       0       1       1       0       0       0       0       0       0       1       0       1       0       0       1       2       0       1       0       1       0       0       0       0       0       0       0       0       0
71      Daucus carota L.                                0       0       0       0       0.1     0       0       0.1     1       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
72      Deschampsia refracta (Lag.) Roemer & Schultz    0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     2       1       0       0       0       0       0       0.1     0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       1       0       0       2       0       0       0       0       3       2       0       0       0       1       0       0       0       0       0       0       0       1       0       2       0       0       0       3       1       0       0       0       0       0       0
73      Dianthus deltoides L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
74      Dianthus scaber Chaix subsp. toletanus          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
75      Digitalis purpurea L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
76      Dipsacus fullonum L.                            0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
77      Eleocharis palustris (L.) Roem. & Schult.       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       2       0       0       1       0       0       1       0       5       0       1       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       4       4       3       0       0       0       0       0       0       0       0.1     0       0       0       0       5       0       0       0       0       0       0       0       0
78      Eleocharis quinqueflora (F.X.Hartm.) O.Schwarz  0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
79      Eleocharis uniglumis (Link) Schultes            0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
80      Elymus caninus (L.) L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0
81      Epilobium hirsutum L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
82      Epilobium obscurum Schreber                     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       1       1       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       1       0       1       1       1       1       0       0       0       0       0       0       0       0       1       0       0.1     1       1       1       0       0       0       0       0       0       0       0       0       2       1       1       1       1
83      Epilobium palustre L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
84      Equisetum palustre L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
85      Equisetum ramosissimum Desf.                    0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0
86      Erica arborea L.                                0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0.1     0       0       0       0       0       0       0       0       0
87      Erica tetralix L.                               0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
88      Eryngium campestre L.                           0       0       0.1     0       0       0.1     0.1     0       0.1     1       1       1       0.1     0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       1       1       0       0.1     0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
89      Festuca ampla Hackel                            0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
90      Festuca arundinacea Schreber                    0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0.1     0       0       0       0       0       0       0       0       0       0
91      Festuca costei (St.Yves) Markgr.-Dannemb.       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       2       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
92      Festuca iberica (Hackel) K. Richter             0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       4       3       0       0       3       1       0       0       0       0       0       0       0       3       3       1       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       2       4       4       2       2       1       1       3       0       1       0       2       1       0       0       0       0       0       0       0       0       0       0       1       2       2       2       0       1       0       0       0       0       0       0       0       0       0.1     0.1     0       0.1     0       1
93      Festuca rothmaleri (Litard.) Markgr.-Dannemb.   0       0       0       3       1       3       2       3       2       1       2       1       2       1       4       1       1       0       0       0       1       1       0       0       1       0       1       2       1       0.1     1       0       0       0       0       0       0       0       0       0       0.1     0       0       2       1       0       4       0       2       0       0       0       0       0       0       0       0       1       0       2       0       0       0       0       1       1       0       0       0       0       0       0       0       0       0       0       0       0       1       1       1       0       0       0       0       0.1     0       0       0       0       0       0
94      Filipendula ulmaria (L.) Maxim.                 0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
95      Foeniculum vulgare Miller                       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
96      Frangula alnus Miller                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     1       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0
97      Fraxinus angustifolia Vahl                      0       0       0       0       1       0       2       3       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0.1     1       0       0       0       5       0.1     0       0       0       0       0       0
98      Galium palustre L.                              0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       1       2       0       0       1       0       0       0       0.1     0       0       0       1       2       1       1       1       1       1       1       0       0       1       0       0       0       0       0       0       0       0       1       0       1       0       2       0       1       1       2       3       0       1       1       2       1       0       0       0       0       0       0       0       0       0       0       0.1     0.1     0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1
99      Galium verum L.                                 0       0       0       0       0.1     0.1     0       0       0       1       0       0       0       0       0       0.1     0.1     0       0       0       0       0       0       0       1       1       1       1       0       0.1     0       0       2       0       0       0       0       0       0       0       0       0       0       0       0.1     0       3       0       0       0       1       0       0       0       0       0       0       1       0       0.1     0       0       0       0       1       0.1     0       0       0       0       0       0       0       1       0       0       0       0       1       0       0       0       1       0       0       0       0       0       0       0       0       0
100     Geum urbanum L.                                 0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0
101     Gladiolus illyricus Koch                        0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
102     Glyceria declinata Bréb.                        0       0       0       0       0       0       0       0       0       0       0       0       0       2       2       0       0       0       2       0       0       0       2       2       0       0       0       0.1     4       4       2       0       0       0       3       0       2       0       1       0       0       1       2       0       2       0       0       0       0       3       1       0       1       0       0       2       0       0       2       0       1       1       3       1       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       2       0       1       0       0       4       2       0       0       0       0       1       2
103     Hedera helix L.                                 0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
105     Heracleum sphondylium L.                        0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
106     Herniaria glabra L.                             0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
107     Hieracium pilosella L.                          0       0       0       0       0       0       0       0       0       2       1       1       1       0       0.1     2       2       0       0       0       0       1       0       0       0       0       0.1     1       0       0       0       0       1       0.1     0       1       1       0       0       0       0       0       0       3       0       3       0       0.1     1       0       1       0.1     0.1     0       0       0       0       0       0       1       0.1     0       0       0       3       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
108     Holcus lanatus L.                               0       0       0       0       0       0       0       2       0       0       0       1       1       1       0       2       1       0       0       3       0       0       0       0       3       0       3       0       3       2       3       2       1       1       1       1       0       1       1       0       2       0       0       0       0       2       0       2       0       0       0       0       0       0       0       3       2       2       2       1       2       0       0       0       1       0       0       0       0       0       0       1       2       1       2       1       2       2       2       1       3       0       0       0       1       1       0       0       0       0       0       0
109     Holcus mollis L.                                0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       3       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       2       0       0       1       0       0       0       0       0       0
110     Hordeum secalinum Schreb.                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
111     Hypericum humifusum L.                          0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
112     Hypericum undulatum Shous.ex Willd.             0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       1       0       0.1     0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0.1     0       0       0       1       1       2       1       0       0       0       0       0       0       0       0       1       0       0       1       0.1     4       0       0       0       1       1       0       0       0       0       0       0       0       0       0
113     Hypochoeris radicata L.                         0       1       1       1       0.1     2       0.1     0       0       0.1     1       1       1       0.1     1       1       2       0       0       0       0.1     0.1     0       0       0       0.1     0.1     0.1     0       0       0       0       1       0.1     0       0.1     0       1       0       0       0.1     1       0       2       1       0       0.1     1       1       0       0       0       0       0       0       1       0       1       0       1       0.1     0       0       0       1       0.1     2       0       0       0       0       0       0       0       0       0       0       0.1     1       0       1       1       1       0       0       0.1     0       0       0       0       0       0
114     Ilex aquifolium L.                              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
116     Jasione montana L.                              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       1       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
117     Juncus acutiflorus Ehrh. ex Hoffm.              0       0       0       0       0       0       0       1       0       2       3       2       2       3       0       1       2       0       0       0       0       1       1       1       2       0       1       2       4       4       5       0       0       2       4       0       0       0       2       0       3       1       4       0       4       0       1       1       0       0       0       0       0       0       0       2       3       1       0       4       4       0       0       0       1       2       3       0       0       0       2       0       2       2       1       1       3       1       1       1       3       1       1       0       0       4       0       0       0       2       2       2
118     Juncus articulatus L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       4       0       0       0       0       0       0       0       0       0       0       1       1       1       2       1       0       0       0       0       0       0       0       0       0       0       1       1       0       1       1       1       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       3       2       2       3       2
119     Juncus bulbosus L.                              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       1       2       0       0
120     Juncus compressus Jacq.                         0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
121     Juncus effusus L.                               0       0       0       0       0       0       0       0       0       0       0       1       0       1       0       0       0       0       1       2       0       0       0       0       0       0       1       0       1       2       1       1       0       1       1       0       0       0       0       2       1       3       0       0       0       0       0       2       0       1       1       0.1     0       2       1       4       1       2       1       0       1       0       1       0       0       3       0       0       0       0       0       0       1       1       3       4       2       1       2       5       0       3       1       0       0       1       0       0       0       0       0       0
122     Juncus heterophyllus Dufour                     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
123     Juncus inflexus L.                              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       2       0       0       0       0       0       0       0
124     Juncus squarrosus L.                            0       0       0       0       0       4       0       0       0       0       0       0       0       0       0       0       0       0       2       3       0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       0       0       0       0       0       0.1     2       0       0       2       0       2       0.1     0       3       0       0       1       1       2       0       1       1       0       1       0       0       0       0       2       1       0       0       0       0       0       0       0.1     0       0       0.1     0       0       1       0.1     1       0       1       0       0       0       1       0       1       1       0       0
125     Juniperus communis L. subsp. nana syme          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       1       0       0       0       0       0       0       0       0.1     0       0.1     0.1     0.1     0.1     0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0.1     0       0       0       0.1     0.1     0.1     0.1     0       0
126     Knautia arvensis (L.) Coulter (Briq.) Szabó     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
127     Lavandula stoechas L. subsp. pedunculata (Miller) Samp. ex Rozeira      0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0
128     Leontodon carpetanus Lange                      0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     1       1       0       0       1       1       1       1       1       2       0       0       1       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       1       0       0       1       0.1     0       0       0       3       2       2       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
129     Leontodon tuberosus L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0
130     Lepidium heterophyllum Bentham                  0       0       0       0       0       0       0       1       0       0.1     0.1     0.1     0.1     0.1     0       0       0       0       0       0       0       0       0.1     0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
131     Linum vienne Miller                             0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
132     Lolium perenne L.                               0       0       0       0       0.1     0       1       1       0.1     0       0       0       0.1     0       3       0       0       0       0       0       2       0.1     0       0       0       0       0       0       0       0.1     0       0       0       0       0       0.1     0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0
133     Lonicera periclymenum L.                        0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
134     Lotus corniculatus L.                           0       0       0.1     0       0       1       1       0.1     0       1       1       1       1       1       0       1       1       0       0       1       0       0       1       1       2       0       2       1       1       1       1       3       2       2       1       0       0       1       1       0       1       2       0       0       0       1       1       0       0       0       0.1     0.1     0.1     0       0       3       1       3       1       2       2       2       1       1       1       0.1     0       0       0       0       0       2       1       1       1       1       1       1       2       1       1       1       1       0       0       0       0       0       0       1       0       3
135     Luzula campestris (L.) DC.                      0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       0       0       0       2       0       2       0       0       0       0       2       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       1       1       1       0       0       1       1       0       0       1       0       1       2       1       1       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0
136     Luzula forsteri (Sm.) DC.                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
137     Lychnis flos-cuculi L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
138     Lepidotis inundata (L.) C. Börner               0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0
139     Lycopus europaeus L.                            0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
140     Lygos sphaerocarpa (L.) Heyw.                   0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
141     Lysimachia vulgaris L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       4       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
142     Lythrum salicaria L.                            0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0
143     Melica ciliata L. ssp. magnolii (Gren.&Godron)Husnot     1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
144     Mentha cervina L.                               0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       4       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
145     Mentha pulegium L.                              0       1       0.1     0       0       0       0       1       0       1       2       2       1       3       3       1       2       2       0       0       0.1     0       0       0       0       0.1     0       1       1       2       2       0       0       2       2       1       2       2       2       0       0       0       0       1       1       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       1       0       0       0       2       3       2       0       0       0       0       0       0       0       0       0       0       0       2       3       0       0       0       0       0       1       0       0       0       0       0       0
146     Mentha suaveolens Ehrh.                         0       0       1       1       0       0       0       1       0       0       0       0.1     0       0       0       0       1       0       0       0       0       0       0       1       2       0       2       0       0       1       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0
147     Merendera pyrenaica (Pourret) P. Four.          0       0       0.1     0       0       0       0.1     0       0       2       1       0       1       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0.1     0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
148     Molinia caerulea (L.) Moench                    0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
149     Myosotis stolonifera (DC) Gay ex Leresche & Levier      0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       1       1       0       0       0.1     0       1       0       1       0       0       1       1       0       1       0       1       0       0       0       2       0       0       0       0       0       0       1       1       0       1       0       0       1       1       0       1       0       0       1       2       0       0       0       0       0       0       0       0       0       0       0       1       1       1       0       0       0       0       0       0       0       0       0       1       1       1       1       1       1
151     Narcissus bulbocodium L.                        0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0.1     0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0.1     0       0
152     Nardus stricta L.                               0       0       0       0       0       0       0       0       0       3       2       2       2       1       0       4       3       0       1       2       0       0       1       1       1       0       4       2       1       1       0       4       3       3       2       4       3       4       0       0       0       4       0       0       1       0       0       4       0       3       2       5       3       5       3       0       3       0       0       4       3       0       0       0       3       3       0       0       0       0       0       0       0       0       2       0.1     1       1       1       0       0       0       1       0       0       0       4       3       2       4       1       3
153     Nepeta nepetella L.                             0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0
154     Oenanthe crocata L.                             0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
155     Ononis repens L.                                0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
156     Orchis coriophora L. subsp. fragrans (Pollini) Sudre    0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
157     Oxalis acetosella L.                            0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
158     Parnassia palustris L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       2       2       1       0
159     Paronychia argentea Lam.                        0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
160     Pedicularis sylvatica L.                        0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       1       0       0       1       0       0       1       0       0       0       0       0       0       0       1       1       1       0       1       1       0.1     1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       1       0       0       0       0       0       0       0.1     0       0
161     Peucedanum officinale L:                        0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
162     Phleum pratense L. subsp. bertoloni (DC.) Bornm.        0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0
163     Pinus sylvestris L.                             0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       3       0       0       0       0       1       0       0       0       0       0       0.1     0       0.1     0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
164     Plantago alpina L. subsp. penyalarensis (Pau) Rivas Martinez    0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0
165     Plantago coronopus L:                           0       0.1     0       0       0       0       0       0       0       0       0       0       0.1     0.1     0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
166     Plantago lanceolata L.                          1       1       1       1       1       1       0       1       1       2       1       1       1       1       1       1       1       0       0.1     0       1       1       1       0       0       0       1       1       0       0       0       0       1       0       0       0       0       0       0       0       0.1     0       0       2       3       2       1       1       2       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0.1     0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0
167     Plantago major L.                               0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       1       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
168     Poa bulbosa L.                                  0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
169     Poa trivialis L.                                0       0       0       0       0       0       1       1       1       0       0       0       0.1     1       1       0       0       0       1       1       1       2       2       2       1       0       0       0       1       1       1       1       0       0       0.1     2       3       1       0       3       3       1       0       0       0       0       0       0       0       0       2       0       0       0       0       3       0       2       3       0       0       3       0       0.1     0       1       1       0       0       0       0       0.1     1       2       2       0       1       1       0       2       2       1       0       0       0       1       0       0       0       0       0       0
170     Polygala vulgaris L.                            0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0
171     Polygonum bistorta L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
172     Potamogeton perfoliatus L.                      0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
173     Potentilla erecta (L.) Räuschel                 0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       1       0       0       0       0       1       0       1       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       3       0.1     1       1       1       1       0       1       0       0       0       3       0       1       1       0       0       0       0       0       0       0       0       3       1       0       0       0       0       0       0       0       0       1       0       0       0       0       1       1       1       0.1     0.1
174     Potentilla hirta L.                             0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
175     Potentilla reptans L.                           0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
176     Prunella grandiflora (L.) Scholler              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
177     Prunella laciniata (L.) L.                      0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       1       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
178     Prunella vulgaris L.                            0       0       0       0       0       0       0       2       0       0.1     0       1       1       1       0       2       1       0       1       1       0       0       1       1       1       0       1       1       1       1       2       0       0       2       1       1       1       1       0       0       1       1       0       0       0       0       0       0       0       0       0.1     0       0       1       0       1       1       1       2       0       2       1       0.1     1       0       1       0       0       0       0       0       0       0       0.1     1       1       1       1       0       0       0       1       1       0       0       0       0       0       0       0       0       0
179     Prunus spinosa L.                               0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0
180     Pteridium aquilinum (L.) Kuhn.                  0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       4       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0
181     Quercus pyrenaica Willd.                        0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
182     Quercus ilex L. subsp. rotundifolia (Lam.) Tab. Morais  0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
183     Ranunculus acris L.                             0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
184     Ranunculus bulbosus L.                          0       1       0.1     1       0       1       1       1       0       1       1       0       1       1       1       1       1       0       3       3       0       0.1     0.1     0       1       0       0       1       0       0       0       0       1       0       0       3       0       2       0       0       1       0       0       0       0       0       1       1       1       1       1       1       0       1       0       2       1       1       0       1       1       0       0       0       1       1       1       0       0       0       0       0       0       1       0       0       0       1       1       0       1       0.1     1       0       0       2       0       1       0       1       0       0.1
185     Ranunculus flammula L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       3       0       0       1       0       0       1       1       0       0       0       0       0       0       0       2       1       0       0       0       0       1       2       0       3       0       1       0       1       0       0       0       0       0       0       0       0       0       0       0       0.1     2       2       0       0       1       0       2       0       0       0       1       0       0       0       1       0       1
186     Ranunculus paludosus Desf.                      0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
187     Ranunculus repens L.                            0       0       0       0       0       0       0       0       0       0       0       0.1     1       1       0       0       0.1     0       0       0       0       0       3       2       0       0       0       0       2       1       1       0       0       0       3       1       3       0       0       4       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       4       0       0       3       4       2       0       0       0       0       0       0       0       0       0       0       3       0       1       0       0       2       0       1       0       0       0       0       0       0       0       0       0       0
188     Rhamnus cathartica L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       4       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
189     Rhinanthus minor L.                             0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
191     Rubus ulmifolius Schott                         0       0       0       0       2       0       0       2       0       0       0       1       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0.1     0       1       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       2       2       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       1       0       0       0       0       0       0       0       0       0
192     Rumex acetosella L.                             0       0       0       0       0       0       0       0.1     1       0.1     1       1       1       0       0       0.1     0.1     0       0       0       0       0       1       1       0       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       1       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
194     Rumex conglomeratus Murray                      0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       1       0       0       1       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0
195     Rumex crispus L.                                0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       2       0       0       0       0       0       0       0
196     Rumex papillaris Boiss. & Reuter                0.1     0       0       1       1       1       0       0.1     1       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       0       0       0       0       0       0       0       0       0       1       0       1       0       0       0       0       0.1     0       0       0       0       0       0       0       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
197     Rumex pulcher L.                                1       0       0.1     0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
198     Sagina nevadensis Boiss. & Reuter               0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0
199     Salix atrocinerea Brot.                         0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       4       0       0       0       0       0       0       0       0       0       1       0       0       0       0.1     0       0       0       1       1       0       0       0       0       0       0       0       0       0
200     Sanguisorba minor Scop. subsp. minor            2       0       0       1       0       0       0       0       0       0.1     0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
201     Sanguisorba officinalis L:                      0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
202     Santolina rosmarinifolia L.                     0       0       0       0       0       0       0       0       0       0.1     0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
203     Scirpus holoschoenus L.                         3       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
204     Scirpus lacustris L.                            0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       5       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
205     Scirpus setaceus L.                             0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     2       0       1       1       0.1     0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0.1     0       0       0       0       1       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0
206     Scolymus hispanicus L.                          1       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
207     Scrophularia auriculata L.                      0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
208     Senecio jacobaea L.                             1       0       0       1       1       0.1     0       2       1       1       1       0.1     2       1       1       1       1       0.1     0       0       1       1       1       0       0       0.1     0       1       0.1     0.1     0       0       0       0       0.1     0       0.1     1       0       0       0       0       0       2       1       3       0       0       0       0       1       0       0       0       0       1       0.1     0       0       0       0       0       0       0       1       1       1       0       0       0       0       2       0       0       0       0       0       0       0.1     0       0       0.1     0       0       0       0.1     0       0       0       0       0       0
209     Silene vulgaris (Moench) Garcke                 0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
210     Sisymbrella aspera (L.) Spach                   0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
211     Spiranthes aestivalis (Poiret) L.C.M. Richard   0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
212     Stellaria graminea L.                           0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
213     Succisa pratensis Moench                        0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       4       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
214     Tamus communis L.                               0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
215     Taraxacum officinale group.                     0       0       0       0       0.1     0       0       0       0       0.1     0       0       0       0       0       0.1     0       0       0.1     0       0       0       1       0.1     0       0       0       0       0       0       0       0       0       0       0       1       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       1       0       0       0       0       0.1     1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
216     Teucrium scorodonia L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
217     Thapsia villosa L.                              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
218     Trifolium fragiferum L.                         0       1       0       0       0       0       0       0       0       0       0       0       2       1       0       0       0       0       0       0       4       4       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       0       0       0       0       0       0       3       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       2       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0
219     Trifolium pratense L.                           0       0       0       0       1       0       0       1       0       0       0       0       0       0.1     0.1     2       1       0       2       1       0       1       1       0.1     0       0       1       1       0       0.1     0       0.1     1       1       0       1       3       0       0       0       1       2       0       0       0       0       0       0       0       2       2       0       0       0       0       1       0       2       1       0       2       1       0       3       0.1     2       1       0       0       0       0       2       0       3       1       1       1       0       1       0       1       0       1       0       0       0       0       0       0       0       0       0
220     Trifolium repens L.                             0       0       0       0       0.1     0       0       1       0       3       2       3       1       2       0       2       2       0       0       0       0       0       2       3       0       0       0       4       1       1       1       0       0       1       2       3       1       2       2       0       0       1       0       0       0       0       0       0       0       0       2       2       1       0       0       2       2       1       0       3       2       3       0       0       2       0       0       0       0       0       0       3       0       0       2       0       0       0       4       2       3       1       2       0       0       0       0       0       0       0       0       1
221     Trisetum flavescens (L.) Beauv.                 0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
222     Urtica dioica L.                                0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
223     Valeriana officinalis L.                        0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
224     Verbascum sinuatum L.                           0       0       0       0       0.1     0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
225     Verbena officinalis L.                          0       0       0       1       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
226     Veronica anagallis-aquatica L.                  0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
227     Veronica catenata Pennell                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       1       0       0       1       1       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
228     Veronica beccabunga L.                          0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
229     Veronica officinalis L.                         0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
230     Veronica scutellata L.                          0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       2       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
231     Veronica serpyllifolia L.                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       1       0       0
232     Viburnum opulus L.                              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
233     Viola canina L.                                 0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
234     Viola odorata L.                                0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
235     Viola palustris L. subsp. Juressi (Link ex K. Wein) Coutinho    0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       0       0       1       0       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       3       1       0       0       0       0
236     Viola riviniana Reichenb.                       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1     0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0
237     Wahlenbergia hederacea (L.) Reich.              0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       0       1       1       0       1       0       0       1       0       0       0       1       0       0       0       0       0       0       0       0       0       0       0       0       1       1       1       0       0       0       0       0       0       0       0       0       0       0       0       0       0.1

Appendix 2. Ecological information for the 232 species. Abbreviations: A1, Axis 1, etc.; min., minimum; max, maximum; len., length; envir. vol., environmental volume; Sites, no. sites found; ab., abundance; Occ., habitat occupancy; Grp., habitat group; Eur., European range; N. Afr., North African range; Cont., continental range; Iber. (km2), Iberian range.

A tab-delimited version of this appendix is available as a text file.

Plant   A1 max  A1 min  A2 max  A2 min  A3 max  A3 min  A1 len. A2 len. A3 len. Envir. vol.  Sites   Mean ab.  Max ab.    Occ. Grp     Eur.  N. Afr.  Cont.    Iber. (km2)
1       0.1645  0.0806  0.1733  0.1301  0.0029  -0.0587 0.0839  0.0432  0.0616  0.0005325160    2       0.55    1       1.0000  2       34      0       34      24780.53
2       0.2037  0.0044  0.1602  0.1094  0.1070  -0.1120 0.1993  0.0508  0.2190  0.0052883640    2       0.55    1       0.4000  4       2       1       3
3       -0.0813 -0.0813 0.0143  0.0143  -0.0425 -0.0425 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  2       35      2       37      541153.51
4       0.4230  -0.1432 0.1878  -0.4377 0.2484  -0.1797 0.5662  0.6255  0.4281  0.3616168164    32      1.54    4       0.5517  1       30      0       30      304092.38
5       0.0044  0.0044  0.1602  0.1602  0.1070  0.1070  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  2       31      2       33      544801.38
6       0.2632  -0.3533 0.3545  -0.4377 0.2224  -0.3766 0.6165  0.7922  0.5990  0.6977517798    62      2.12    5       0.7848  3       10
7       0.2037  -0.4238 0.1878  -0.1812 0.1041  -0.1797 0.6275  0.3690  0.2838  0.1567323694    12      1.10    2       0.2308  1       38
8       0.0806  0.0395  0.1301  0.1106  0.0444  0.0029  0.0411  0.0195  0.0415  0.0000793288    2       0.55    1       1.0000  2       21      0       21      8369.09
9       -0.2885 -0.2885 0.0514  0.0514  0.0333  0.0333  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  2       31      6       37      256809.35
10      -0.3396 -0.3526 -0.0592 -0.0961 0.0883  0.0704  0.0130  0.0369  0.0179  0.0000204800    2       0.10    0.1     1.0000  4       28      0       28      1562.88
11      -0.2724 -0.2992 -0.2719 -0.3640 -0.3119 -0.3766 0.0268  0.0921  0.0647  0.0003808947    2       0.10    0.1     1.0000  4       25                      30166.00
12      0.0855  0.0855  0.1873  0.1873  -0.0596 -0.0596 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  1       34      4       38      187313.96
13      0.2101  -0.3533 0.2071  -0.1800 0.2181  -0.2050 0.5634  0.3871  0.4231  0.2200843953    10      0.74    2       0.1613  1       20      1       21      53635.63
14      -0.2659 -0.3323 -0.0229 -0.0579 0.2181  0.1041  0.0664  0.0350  0.1140  0.0006318983    2       1.05    2       1.0000  1       28      0       28
15      0.2377  0.0450  0.0869  -0.1812 -0.0372 -0.1797 0.1927  0.2681  0.1425  0.0175589930    7       1.57    3       0.4375  1       17      3       20      423466.32
16      -0.2843 -0.2843 0.0401  0.0401  -0.0682 -0.0682 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       6                       275843.53
17      0.4230  -0.1432 0.2415  -0.1881 0.2224  -0.1797 0.5662  0.4296  0.4021  0.2332783433    33      1.10    2       0.6000  3       38
18      0.2037  -0.1432 0.1094  -0.1812 0.0062  -0.1797 0.3469  0.2906  0.1859  0.0446977345    4       0.55    1       0.1905  1       17                      561793.72
19      -0.2885 -0.2885 0.0514  0.0514  0.0333  0.0333  0.0000  0.0000  0.0000  0.0000000000    1       2.00    2       1.0000  4       4       3       7       371298.95
20      0.0395  -0.2885 0.1106  0.0221  0.0444  -0.1832 0.3280  0.0885  0.2276  0.0157578000    5       0.84    2       0.5000  4       2
21      0.0044  0.0044  0.1602  0.1602  0.1070  0.1070  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  2       35      2       37      573312.31
22      -0.0615 -0.3387 0.3545  0.0531  0.1412  -0.0643 0.2772  0.3014  0.2055  0.0409500571    3       0.70    1       0.3000  4       9
23      -0.0955 -0.0955 -0.0792 -0.0792 -0.0060 -0.0060 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  1       19      3       22      150475.96
24      0.2283  -0.2885 0.3545  -0.1800 0.0333  -0.2050 0.5168  0.5345  0.2383  0.1570002950    14      1.38    4       0.3111  3       35      2       37      407959.35
25      0.2130  -0.1432 0.2105  0.1086  0.1452  -0.1115 0.3562  0.1019  0.2567  0.0222228717    5       1.60    4       0.4167  2       26      0       26      108340.66
26      0.1574  -0.2247 0.1873  -0.1361 -0.0425 -0.1758 0.3821  0.3234  0.1333  0.0392874114    5       1.42    3       0.3571  3       35      4       39      562445.76
27      0.2377  -0.1432 0.1602  -0.4377 0.2224  -0.1797 0.3809  0.5979  0.4021  0.2184136672    21      0.83    2       0.4773  3       3       1       4       397175.46
28      -0.2885 -0.2885 0.0514  0.0514  0.0333  0.0333  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       24      3       27      527180.13
29      0.2757  -0.0813 0.1438  -0.1036 0.0662  -0.0425 0.3570  0.2474  0.1087  0.0228983225    3       1.37    3       0.2500  2       30      2       32      358628.19
30      -0.2885 -0.2885 0.0514  0.0514  0.0333  0.0333  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  3       26      6       32      418306.13
31      0.2130  0.0278  0.2105  -0.0330 0.1452  -0.1115 0.1852  0.2435  0.2567  0.0276103574    3       1.37    3       0.2000  1       30      1       31      116939.62
32      -0.4238 -0.4238 -0.0139 -0.0139 0.0626  0.0626  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       1                       148519.83
33      0.2632  -0.1964 0.2415  -0.0501 0.0441  -0.1132 0.4596  0.2916  0.1573  0.0502808341    4       0.10    0.1     0.1818  4       2
34      0.3388  -0.2593 0.2415  -0.4377 0.2224  -0.1832 0.5981  0.6792  0.4056  0.3929846956    36      1.38    4       0.5070  1       9       1       10      79273.85
35      0.4230  0.0450  0.1733  -0.2314 0.1976  -0.1797 0.3780  0.4047  0.3773  0.1376632508    25      1.41    3       0.6250  1       20      0       21
36      -0.1722 -0.3924 0.2071  -0.3640 0.1324  -0.3766 0.2202  0.5711  0.5090  0.1526699167    12      2.00    3       0.5455  3       21                      486663.58
37      0.2127  -0.3396 0.0514  -0.0592 0.0704  -0.1132 0.5523  0.1106  0.1836  0.0267490929    4       0.55    1       0.2857  3       29      7       36      481587.06
38      0.4230  0.0278  0.2415  -0.2314 0.2484  -0.1797 0.3952  0.4729  0.4281  0.1908260626    24      1.63    4       0.5106  1       33      4       37      215077.55
39      0.2283  -0.0813 0.1301  -0.4377 0.1503  -0.1727 0.3096  0.5678  0.3230  0.1354269426    10      0.83    2       0.3125  1       36      6       42      499283.53
40      0.1574  -0.3087 0.1878  -0.1361 0.1463  -0.1758 0.4661  0.3239  0.3221  0.1159810369    9       0.70    1       0.2195  3       31      3       34      213631.31
41      0.0214  -0.4238 0.0686  -0.1361 0.1463  -0.0916 0.4452  0.2047  0.2379  0.0517098945    9       0.40    1       0.4286  4       31      4       35      427350.03
42      0.4230  0.0746  0.1321  -0.2314 0.2484  -0.1797 0.3484  0.3635  0.4281  0.1293105625    20      2.05    4       0.5556  1       33      4       37      286919.74
43      0.3388  -0.3533 0.2415  -0.4377 0.2224  -0.1779 0.6921  0.6792  0.4003  0.4488056629    37      1.30    3       0.4684  1       32      6       38      372490.06
44      0.2757  0.0044  0.1602  -0.2438 0.2224  -0.1797 0.2713  0.4040  0.4021  0.1051166335    12      1.84    3       0.3243  1       32      1       33      52781.50
45      -0.2308 -0.2314 -0.0740 -0.1706 -0.0628 -0.1177 0.0006  0.0966  0.0549  0.0000075894    2       0.10    0.1     1.0000  4       14                      427522.68
46      -0.2362 -0.2838 0.2423  0.0593  0.0702  -0.0049 0.0476  0.1830  0.0751  0.0015602859    2       0.10    0.1     0.6667  4       3
47      0.4230  -0.3533 0.2415  -0.4377 0.2224  -0.3766 0.7763  0.6792  0.5990  0.7532866960    51      1.66    5       0.5930  1       7                       238655.27
48      -0.1727 -0.1727 0.3545  0.3545  -0.0643 -0.0643 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       5                       70939.55
49      0.0249  -0.1864 0.1878  0.0143  0.0583  -0.1779 0.2113  0.1735  0.2362  0.0206530921    4       1.78    3       0.6667  4       16      0       16      143997.94
50      -0.4238 -0.4238 -0.0139 -0.0139 0.0626  0.0626  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       26      0       16      156483.78
51      0.3395  -0.2247 0.2415  -0.1361 0.1278  -0.1388 0.5642  0.3776  0.2666  0.1354663483    21      0.44    1       0.4565  1       38      3       41      229492.82
52      0.2130  0.2130  0.2105  0.2105  -0.1115 -0.1115 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  1       17                      102288.66
53      0.1300  -0.3533 0.0834  -0.1706 0.0187  -0.1727 0.4833  0.2540  0.1914  0.0560400684    9       1.46    2       0.4091  3       17                      115638.72
54      0.2127  -0.4238 0.1086  -0.1361 0.0626  -0.1132 0.6365  0.2447  0.1758  0.0653066580    4       0.55    1       0.1379  1       2                       325991.47
55      0.2283  0.2130  0.2105  0.0636  -0.1084 -0.1115 0.0153  0.1469  0.0031  0.0000166181    2       0.55    1       1.0000  1       27      1       28      580962.99
56      0.1439  -0.0813 0.0143  0.0023  -0.0425 -0.1758 0.2252  0.0120  0.1333  0.0008591836    2       0.10    0.1     1.0000  3       35      7       42      279519.09
57      -0.0244 -0.3396 0.2423  -0.1706 0.1412  -0.1247 0.3152  0.4129  0.2659  0.0825383230    12      1.27    4       0.5000  4       36      10      46      491671.51
58      -0.0818 -0.0818 -0.0733 -0.0733 -0.0647 -0.0647 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       20      2       22      536938.42
59      0.0958  -0.2659 0.1086  -0.1800 0.1041  -0.2050 0.3617  0.2886  0.3091  0.0769573407    6       0.25    1       0.2308  3       32                      166107.18
60      -0.3387 -0.3387 0.0733  0.0733  0.1412  0.1412  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       19                      142726.66
61      0.3395  0.1651  0.1321  0.0536  0.1278  -0.0284 0.1744  0.0785  0.1562  0.0051003899    3       0.40    1       0.7500  4       19
62      0.0249  -0.2247 0.1878  -0.1361 -0.0477 -0.0690 0.2496  0.3239  0.0213  0.0041071574    2       0.55    1       0.6667  3       24      2       26      191694.30
63      -0.0818 -0.4238 0.2423  -0.3640 0.1463  -0.3766 0.3420  0.6063  0.5229  0.2586059588    15      2.47    4       0.5000  3       27
64      0.2632  -0.4238 0.2423  -0.4377 0.2181  -0.3766 0.6870  0.6800  0.5947  0.6626280249    55      1.73    4       0.6875  3       34      1       35
65      0.2037  -0.4238 0.2423  -0.1361 0.0702  -0.1120 0.6275  0.3784  0.1822  0.1031856827    9       1.36    3       0.2368  1       22                      25087.04
66      0.1645  0.1645  0.1733  0.1733  -0.0587 -0.0587 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       3       2       5       282151.44
67      0.2130  0.2130  0.2105  0.2105  -0.1115 -0.1115 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  3       25      1       26      52131.18
68      0.1439  -0.2885 0.2423  -0.1800 0.1070  -0.2050 0.4324  0.4223  0.3120  0.1358837652    9       0.60    1       0.2093  4       38      10      48      580963.00
69      0.0806  -0.1864 0.1602  0.1009  0.1070  -0.1779 0.2670  0.0593  0.2849  0.0107588194    3       0.40    1       0.6000  3       5
70      0.2377  -0.3087 0.2071  -0.4377 0.2224  -0.1779 0.5464  0.6448  0.4003  0.3363779512    31      1.05    3       0.4769  3       34      3       37      125725.73
71      -0.0615 -0.4238 0.0531  -0.1800 0.0883  -0.2050 0.3623  0.2331  0.2933  0.0590784214    6       0.40    1       0.3529  4       36      10      46      580962.99
72      0.2101  -0.3533 0.1602  -0.4377 0.2224  -0.1779 0.5634  0.5979  0.4003  0.3216156678    15      1.48    3       0.2542  1       2       2       4       351492.04
73      -0.0615 -0.0615 0.0531  0.0531  0.0583  0.0583  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  3       27      0       27      29810.87
74      0.2101  0.2101  0.1602  0.1602  -0.0017 -0.0017 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       3       0       3
75      0.2632  0.2632  0.2415  0.2415  -0.0092 -0.0092 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       18      1       19      252252.83
76      -0.2843 -0.2843 0.0401  0.0401  -0.0682 -0.0682 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  3       29      5       34      445255.49
77      0.2377  -0.3544 0.1878  -0.1487 0.1870  -0.1577 0.5921  0.3365  0.3447  0.1638051775    13      2.32    5       0.2167  1       38      10      48      580963.00
78      0.2757  0.1302  0.0174  -0.1036 0.0525  -0.0517 0.1455  0.1210  0.1042  0.0043754457    3       0.70    1       1.0000  1       31      3       34      33112.61
79      0.0450  0.0450  0.0029  0.0029  -0.0410 -0.0410 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  1       31      2       33      25116.40
80      0.1574  -0.0813 0.0961  0.0143  -0.0425 -0.1717 0.2387  0.0818  0.1292  0.0060169229    2       1.00    1       0.6667  3       33      0       33      191573.28
81      0.1773  -0.1432 0.1086  0.0348  0.0620  0.0062  0.3205  0.0738  0.0558  0.0031479281    2       1.05    2       0.5000  1       35      9       44      365507.48
82      0.4230  -0.1432 0.2105  -0.2314 0.2224  -0.1132 0.5662  0.4419  0.3356  0.2002728279    21      0.88    2       0.4286  1       30      3       33      120392.99
83      0.2101  0.1685  0.1602  -0.1236 0.0894  -0.0017 0.0416  0.2838  0.0911  0.0025652536    2       1.00    1       0.5000  1       31      1       32      37181.94
84      -0.1432 -0.1432 0.1086  0.1086  0.0062  0.0062  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  1       32      2       34      118250.56
85      -0.0813 -0.0813 0.0143  0.0143  -0.0425 -0.0425 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  1       29      10      39      418302.20
86      0.2130  -0.0813 0.2105  0.0143  0.1452  -0.1115 0.2943  0.1962  0.2567  0.0353525989    4       0.55    1       0.2353  2       14      4       18      333969.13
87      0.3305  0.3305  -0.0456 -0.0456 0.1578  0.1578  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  1       16      0       16      144913.67
88      0.2037  -0.3526 0.3545  -0.1706 0.1412  -0.1727 0.5563  0.5251  0.3139  0.2186999106    15      0.53    2       0.2885  4       29                      559071.81
89      0.1574  -0.2838 0.2423  0.0961  0.0702  -0.1717 0.4412  0.1462  0.2419  0.0372155941    2       2.00    2       0.2000  3       2
90      -0.1722 -0.2992 0.1009  -0.3640 -0.1779 -0.3766 0.1270  0.4649  0.1987  0.0279812651    4       1.53    3       1.0000  1       35      10      45      67689.86
91      0.2236  0.1302  0.1255  -0.0531 -0.0419 -0.1577 0.0934  0.1786  0.1158  0.0046072640    4       1.25    2       0.5714  4       5
92      0.4230  0.0044  0.2415  -0.4377 0.2484  -0.1132 0.4186  0.6792  0.3616  0.2452062494    28      1.87    4       0.6222  4       2
93      0.2037  -0.3526 0.3545  -0.3640 0.1463  -0.3766 0.5563  0.7185  0.5229  0.4984948613    35      1.49    4       0.5556  3       1
94      0.0806  -0.1432 0.1301  0.1086  0.0062  0.0029  0.2238  0.0215  0.0033  0.0000378720    2       1.00    1       1.0000  1       30      0       30      125554.43
95      -0.2362 -0.2362 0.0186  0.0186  -0.1247 -0.1247 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       29                      571691.99
96      0.1651  -0.0813 0.1602  0.0143  0.1452  -0.0425 0.2464  0.1459  0.1877  0.0160940920    4       0.33    1       0.4444  1       30      5       35      263775.76
97      0.0855  -0.3323 0.1873  -0.1800 0.2181  -0.2050 0.4178  0.3673  0.4231  0.1548597668    11      1.57    5       0.2821  3       19      6       25      488712.60
98      0.3395  -0.1432 0.2415  -0.4377 0.2224  -0.1717 0.4827  0.6792  0.3941  0.3081680586    29      1.09    3       0.5273  1       31      6       37      414301.28
99      0.2101  -0.3396 0.1838  -0.0945 0.1463  -0.1758 0.5497  0.2783  0.3221  0.1175265208    21      0.80    3       0.4884  3       33      7       40      306733.06
100     0.1579  -0.2247 0.1086  -0.1881 0.2224  -0.0690 0.3826  0.2967  0.2914  0.0788965969    4       0.55    1       0.2105  2       33      5       38      224035.24
101     -0.3396 -0.3396 -0.0592 -0.0592 0.0704  0.0704  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       13
102     0.3305  -0.3323 0.2415  -0.2438 0.2224  -0.2050 0.6628  0.4853  0.4274  0.3278940382    31      1.91    4       0.4133  1       19      0       19      37883.09
103     -0.1432 -0.2885 0.1086  0.0514  0.0333  0.0062  0.1453  0.0572  0.0271  0.0005372014    2       0.55    1       0.5000  4       33      8       41      527372.32
105     0.0806  0.0806  0.1301  0.1301  0.0029  0.0029  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  2       31      3       34      145551.17
106     0.0452  -0.0818 0.0608  -0.1487 0.0187  -0.0709 0.1270  0.2095  0.0896  0.0056859360    3       2.00    3       0.5000  4       30      8       38      89580.39
107     0.3395  -0.2992 0.3545  -0.3640 0.2224  -0.3766 0.6387  0.7185  0.5990  0.6556268372    25      1.11    3       0.3125  3       33      4       37      215181.71
108     0.2632  -0.4238 0.2423  -0.4377 0.2224  -0.1797 0.6870  0.6800  0.4021  0.4480288024    41      1.73    3       0.5325  3       37      5       42      530951.41
109     0.2130  -0.3087 0.2105  -0.0229 0.1463  -0.1832 0.5217  0.2334  0.3295  0.0956936938    7       1.17    3       0.1842  3       30      1       31      200958.44
110     -0.0965 -0.2724 -0.1800 -0.2719 -0.2050 -0.3119 0.1759  0.0919  0.1069  0.0041215946    2       1.00    1       1.0000  3       19      4       23      86895.95
111     0.0214  0.0214  -0.0047 -0.0047 -0.0916 -0.0916 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  1       22      3       25      222352.64
112     0.2632  -0.1722 0.2415  -0.1881 0.2224  -0.1832 0.4354  0.4296  0.4056  0.1809492783    16      1.08    4       0.3333  3       5       2       7       254180.19
113     0.2632  -0.3533 0.3545  -0.3640 0.2224  -0.3766 0.6165  0.7185  0.5990  0.6328384925    43      0.74    2       0.5513  4       32      4       36      580962.99
114     0.0278  0.0278  0.1327  0.1327  0.1452  0.1452  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  2       22      5       27      124317.73
116     0.2037  -0.2838 0.3545  0.0585  0.0702  -0.1120 0.4875  0.2960  0.1822  0.0627077063    4       0.55    1       0.2353  4       29      3       32      431840.80
117     0.4230  -0.3533 0.2415  -0.3640 0.2224  -0.3766 0.7763  0.6055  0.5990  0.6715475478    50      2.16    5       0.5882  1       25      3       28      259951.36
118     0.4230  0.1251  0.1602  -0.2314 0.2224  -0.1797 0.2979  0.3916  0.4021  0.1118802610    19      1.48    4       0.6129  1       38      5       43      555893.67
119     0.4230  0.0278  0.2105  -0.0788 0.1950  -0.1115 0.3952  0.2893  0.3065  0.0835799409    7       1.43    2       0.2414  1       28      2       30      122006.45
120     -0.4238 -0.4238 -0.0139 -0.0139 0.0626  0.0626  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  1       33      0       33      395689.81
121     0.3395  -0.3087 0.2415  -0.4377 0.2224  -0.1832 0.6482  0.6792  0.4056  0.4259031595    38      1.66    5       0.4935  1       36      6       42      530316.43
122     0.0790  0.0790  -0.0651 -0.0651 0.1557  0.1557  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  1       7
123     -0.1964 -0.3323 0.0221  -0.0579 0.2181  0.0441  0.1359  0.0800  0.1740  0.0045119565    2       1.50    2       0.5000  1       32      7       39      563192.94
124     0.4230  -0.3396 0.2071  -0.4377 0.2484  -0.1779 0.7626  0.6448  0.4263  0.4999693415    28      1.30    4       0.3590  1       20      1       21      45734.24
125     0.4230  -0.0813 0.1438  -0.1848 0.2484  -0.0675 0.5043  0.3286  0.3159  0.1248568473    14      0.23    1       0.4667  4       34      3       37      158188.02
126     0.0806  0.0806  0.1301  0.1301  0.0029  0.0029  0.0000  0.0000  0.0000  0.0000000000    1       2.00    2       1.0000  2       13      0       13      65771.49
127     0.0044  -0.2838 0.2423  0.0186  0.1070  -0.1247 0.2882  0.2237  0.2317  0.0356280625    3       0.40    1       0.2727  4       2       7       9       378753.64
128     0.2377  -0.3533 0.1838  -0.1881 0.2224  -0.1797 0.5910  0.3719  0.4021  0.2107919123    18      1.13    3       0.2769  4       1                       73108.60
129     -0.0813 -0.0813 0.0143  0.0143  -0.0425 -0.0425 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  4       13                      256039.86
130     0.2127  -0.2247 0.0608  -0.1361 -0.0647 -0.1727 0.4374  0.1969  0.1080  0.0221847461    8       0.33    1       0.7273  3       12      0       12      128055.90
131     -0.1999 -0.1999 0.0825  0.0825  0.0232  0.0232  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  3       17      8       25      432279.26
132     0.2236  -0.4238 0.0834  -0.3640 0.1041  -0.3766 0.6474  0.4474  0.4807  0.3320848082    14      0.76    3       0.2857  3       38      8       46      540565.39
133     0.2130  0.2130  0.2105  0.2105  -0.1115 -0.1115 0.0000  0.0000  0.0000  0.0000000000    1       2.00    2       1.0000  3       20      1       21      242340.07
134     0.4230  -0.4238 0.2423  -0.4377 0.2224  -0.1832 0.8468  0.6800  0.4056  0.5570496682    58      1.22    3       0.6667  3       38      5       43      580963.00
135     0.3395  -0.1432 0.2415  -0.4377 0.2224  -0.1758 0.4827  0.6792  0.3982  0.3113740699    18      1.17    2       0.3158  4       33      2       35      571166.00
136     0.2127  0.0855  0.1873  -0.0501 -0.0596 -0.1132 0.1272  0.2374  0.0536  0.0038604580    2       0.55    1       0.5000  4       22
137     0.1773  -0.1432 0.1086  0.0348  0.0620  0.0062  0.3205  0.0738  0.0558  0.0031479281    2       1.50    2       0.5000  2       33      0       33      37954.93
138     0.3094  0.3094  -0.0788 -0.0788 0.1950  0.1950  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  1       25      0       25      451572.03
139     0.0249  0.0249  0.1878  0.1878  -0.0477 -0.0477 0.0000  0.0000  0.0000  0.0000000000    1       2.00    2       1.0000  1       34      6       40      71061.61
140     -0.2362 -0.2362 0.0186  0.0186  -0.1247 -0.1247 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       2       3       5
141     0.0855  0.0855  0.1873  0.1873  -0.0596 -0.0596 0.0000  0.0000  0.0000  0.0000000000    1       4.00    4       1.0000  1       32      2       34      192933.34
142     -0.1864 -0.1864 0.1009  0.1009  -0.1779 -0.1779 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  1       33      6       39      539225.83
143     -0.2362 -0.2362 0.0186  0.0186  -0.1247 -0.1247 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  4       28      6       34      580963.00
144     0.0790  -0.3924 0.0560  -0.0651 0.1557  0.1324  0.4714  0.1211  0.0233  0.0031724578    2       3.50    4       1.0000  1       3       2       5       150882.71
145     0.2377  -0.3533 0.3545  -0.2719 0.2224  -0.3119 0.5910  0.6264  0.5343  0.4717703206    34      1.60    3       0.4474  1       28      9       37      412260.65
146     0.2632  -0.4238 0.2415  -0.1706 0.0626  -0.1758 0.6870  0.4121  0.2384  0.1609799596    13      1.32    3       0.2453  1       26      5       31      573191.79
147     0.3395  -0.2724 0.1838  -0.2719 0.1278  -0.3119 0.6119  0.4557  0.4397  0.2924301580    10      0.56    2       0.1639  4       3
148     0.0249  0.0249  0.1878  0.1878  -0.0477 -0.0477 0.0000  0.0000  0.0000  0.0000000000    1       2.00    2       1.0000  2       30      6       36      221141.94
149     0.4230  0.0278  0.2415  -0.4377 0.2484  -0.1797 0.3952  0.6792  0.4281  0.2740728731    28      1.04    2       0.5714  1       3       0       3
151                                                                             0.3921523124    7       0.23    1       0.1129  3       3
152     0.4230  -0.3087 0.2071  -0.4377 0.2484  -0.1758 0.7317  0.6448  0.4242  0.4773478376    49      2.51    5       0.6622  3       32      2       34      184360.09
153     -0.0813 -0.0813 0.0143  0.0143  -0.0425 -0.0425 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       4       0       4       230442.05
154     0.0249  -0.0965 0.1878  -0.1800 -0.0477 -0.2050 0.1214  0.3678  0.1573  0.0167519491    2       0.55    1       0.5000  3       10
155     -0.2838 -0.2843 0.2423  0.0401  0.0702  -0.0682 0.0005  0.2022  0.1384  0.0000333729    2       0.55    1       1.0000  4       24      8       32      232042.11
156     0.0746  -0.3396 0.0534  -0.0592 0.0704  0.0464  0.4142  0.1126  0.0240  0.0026697214    3       0.10    0.1     1.0000  3       27
157     0.2130  0.2130  0.2105  0.2105  -0.1115 -0.1115 0.0000  0.0000  0.0000  0.0000000000    1       2.00    2       1.0000  2       31      1       32      66522.80
158     0.4230  0.2947  0.0714  -0.2314 0.1976  0.0501  0.1283  0.3028  0.1475  0.0136672381    4       1.75    2       0.6667  1       28      2       30      94431.49
159     0.0214  0.0214  -0.0047 -0.0047 -0.0916 -0.0916 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       10      10      20      328031.09
160     0.4230  -0.0813 0.2415  -0.1812 0.1278  -0.1797 0.5043  0.4227  0.3075  0.1563408784    14      0.81    1       0.2979  2       19      0       19      209846.05
161     -0.2724 -0.2724 -0.2719 -0.2719 -0.3119 -0.3119 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  3       13                      49837.62
162     -0.0615 -0.3533 0.0834  -0.3640 0.2181  -0.3766 0.2918  0.4474  0.5947  0.1851763064    7       0.61    1       0.2414  3       36      5       41      522571.29
163     0.2377  -0.0197 0.2105  -0.2438 0.2224  -0.1132 0.2574  0.4543  0.3356  0.0936007747    9       0.72    3       0.2647  4       31      1       32      52698.75
164     0.2588  0.2588  -0.1848 -0.1848 0.2484  0.2484  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  2       7       0       7       133833.57
165     0.1300  -0.2362 0.3545  -0.1800 0.0573  -0.2050 0.3662  0.5345  0.2623  0.1224533164    7       0.36    1       0.2121  4       26      8       34      493000.41
166     0.2127  -0.4238 0.3545  -0.3640 0.1412  -0.3766 0.6365  0.7185  0.5178  0.5647983744    34      1.04    3       0.5152  4       38      9       47      569172.11
167     0.2236  -0.4238 0.1838  -0.0441 0.0626  -0.1577 0.6474  0.2279  0.2203  0.0775242778    6       1.00    1       0.1818  2       38      10      48      580734.37
168     -0.2362 -0.2362 0.0186  0.0186  -0.1247 -0.1247 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       33      8       41      552179.40
169     0.2632  -0.4238 0.2415  -0.4377 0.1503  -0.3766 0.6870  0.6792  0.5269  0.5863930636    42      1.44    3       0.5676  3       38      9       47      555869.64
170     0.1574  -0.0813 0.1301  0.0143  0.0029  -0.1717 0.2387  0.1158  0.1746  0.0115109569    3       0.10    0.1     0.7500  3       32      1       33      184293.28
171     0.0806  -0.1432 0.1301  0.1086  0.0444  0.0029  0.2238  0.0215  0.0415  0.0004762696    3       1.00    1       1.0000  1       29      2       31      59216.07
172     0.2377  0.1302  0.0869  -0.0531 -0.0419 -0.0481 0.1075  0.1400  0.0062  0.0002225535    2       1.50    2       1.0000  1       32      5       37      31731.36
173     0.4230  -0.1432 0.1602  -0.2314 0.2224  -0.1758 0.5662  0.3916  0.3982  0.2105814033    23      1.19    3       0.4600  2       33      2       35      189706.62
174     -0.2885 -0.2885 0.0514  0.0514  0.0333  0.0333  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  4       4                       10512.01
175     0.1447  -0.2843 0.0401  -0.2438 0.0446  -0.0682 0.4290  0.2839  0.1128  0.0327670992    2       1.50    2       0.1538  4       36      6       42      572893.48
176     0.0806  0.0806  0.1301  0.1301  0.0029  0.0029  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  2       24      0       24      157176.25
177     0.2037  -0.3387 0.3545  0.0593  0.1412  -0.1717 0.5424  0.2952  0.3129  0.1194944704    6       1.03    3       0.3158  3       25      5       30      242980.87
178     0.3388  -0.3087 0.2415  -0.4377 0.2224  -0.1832 0.6475  0.6792  0.4056  0.4254432208    42      1.06    2       0.5526  3       37      6       43      507177.60
179     -0.0813 -0.2885 0.1086  0.0143  0.0333  -0.0425 0.2072  0.0943  0.0758  0.0035324568    3       1.37    3       0.5000  3       33      3       36      238949.93
180     0.2130  -0.0813 0.2105  -0.0501 0.1452  -0.1758 0.2943  0.2606  0.3210  0.0587186304    6       2.02    4       0.2143  4       36      8       44      360637.82
181     0.0855  -0.2885 0.1873  -0.1800 0.0583  -0.2050 0.3740  0.3673  0.2633  0.0862679745    5       0.28    1       0.1613  4       3       1       4       205717.02
182     -0.2362 -0.2362 0.0186  0.0186  -0.1247 -0.1247 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  4       15      4       19      461755.36
183     0.0806  0.0395  0.1301  0.1106  0.0444  0.0029  0.0411  0.0195  0.0415  0.0000793288    2       1.50    2       1.0000  2       32      0       32      92628.34
184     0.4230  -0.3533 0.2415  -0.4377 0.2224  -0.3766 0.7763  0.6792  0.5990  0.7532866960    48      1.09    3       0.5581  3       27      6       33      460455.56
185     0.4230  -0.2659 0.2415  -0.1881 0.2224  -0.1832 0.6889  0.4296  0.4056  0.2863021539    20      1.51    3       0.2857  1       33      3       36      152234.57
186     -0.0818 -0.2843 0.0401  -0.0733 -0.0647 -0.0682 0.2025  0.1134  0.0035  0.0001916957    2       0.55    1       1.0000  4       11      8       19      399745.18
187     0.2632  -0.2098 0.2415  -0.1288 0.1452  -0.1832 0.4730  0.3703  0.3284  0.1371905549    23      1.97    4       0.4792  1       36      3       39      441628.53
188     0.2130  -0.0965 0.2105  -0.1800 -0.1115 -0.2050 0.3095  0.3905  0.0935  0.0269525285    2       2.50    4       0.2000  1       29      3       32      75712.57
189     0.0395  0.0395  0.1106  0.1106  0.0444  0.0444  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  2       31      0       31      70052.18
191     0.2632  -0.2885 0.2415  -0.1361 0.1452  -0.1120 0.5517  0.3776  0.2572  0.1277944948    13      1.18    3       0.2708  3       21                      526110.24
192     0.2283  -0.3526 0.3545  -0.1361 0.0883  -0.1727 0.5809  0.4906  0.2610  0.1774089964    16      0.84    2       0.3077  4       39      4       43      382698.32
194     0.0395  -0.3533 0.1106  -0.1800 0.1041  -0.2050 0.3928  0.2906  0.3091  0.0841535237    7       0.87    1       0.2500  3       33      8       41      534660.79
195     -0.0197 -0.3924 0.0560  -0.1800 0.2181  -0.2050 0.3727  0.2360  0.4231  0.0887606824    6       0.42    2       0.2500  3       37      8       45      566439.62
196     0.0806  -0.3526 0.2423  -0.1361 0.1412  -0.1247 0.4332  0.3784  0.2659  0.1039594862    15      0.70    1       0.4545  4       2       0       2
197     -0.2308 -0.2885 0.0514  -0.1706 0.0333  -0.1247 0.0577  0.2220  0.1580  0.0048271644    3       0.40    1       0.5000  4       23      9       32      572593.12
198     0.3348  0.3348  -0.0166 -0.0166 0.0564  0.0564  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  2       22      2       24      47870.39
199     0.2632  -0.1722 0.2415  0.0143  0.1070  -0.1832 0.4354  0.2272  0.2902  0.0684700069    7       1.73    4       0.2692  1       10                      443192.99
200     0.0452  -0.2843 0.1106  -0.1800 0.0444  -0.2050 0.3295  0.2906  0.2494  0.0569578634    7       0.50    2       0.3043  4       33      10      43      580962.99
201     0.1447  0.1447  -0.2438 -0.2438 0.0446  0.0446  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  2       29      4       33      24004.99
202     0.0455  0.0452  0.0608  -0.0108 -0.0709 -0.0822 0.0003  0.0716  0.0113  0.0000005789    2       0.10    0.1     1.0000  4       3
203     -0.2308 -0.4238 0.2423  -0.1706 0.1463  -0.1247 0.1930  0.4129  0.2710  0.0515083567    5       2.22    3       0.2778  3       26
204     -0.1845 -0.1845 -0.1419 -0.1419 0.1870  0.1870  0.0000  0.0000  0.0000  0.0000000000    2       3.00    5       1.0000  1       35      8       43      552323.36
205     0.2632  -0.3533 0.2415  -0.1812 0.0573  -0.1797 0.6165  0.4227  0.2370  0.1473058133    14      0.88    2       0.2692  3       27      2       29      159174.08
206     -0.2362 -0.4238 0.0186  -0.0961 0.0883  -0.1247 0.1876  0.1147  0.2130  0.0109315581    3       1.00    1       0.7500  4       18                      478763.06
207     -0.1432 -0.1432 0.1086  0.1086  0.0062  0.0062  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  1       16                      494359.92
208     0.2632  -0.4238 0.3545  -0.3640 0.1463  -0.3766 0.6870  0.7185  0.5229  0.6156138230    38      0.87    3       0.5067  4       30      3       33      349750.30
209     -0.2885 -0.2885 0.0514  0.0514  0.0333  0.0333  0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  3       36      9       45      580962.99
210     0.1773  -0.0244 0.0348  -0.1487 0.0620  0.0187  0.2017  0.1835  0.0433  0.0038223995    2       0.55    1       1.0000  3       3       2       5       341149.41
211     0.1496  0.1496  0.0078  0.0078  -0.1388 -0.1388 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  3       13
212     0.0395  0.0395  0.1106  0.1106  0.0444  0.0444  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  2       32      1       33      145816.87
213     0.0806  0.0395  0.1301  0.1106  0.0444  0.0029  0.0411  0.0195  0.0415  0.0000793288    2       2.50    4       1.0000  3       30      3       33      153464.55
214     -0.2885 -0.2885 0.0514  0.0514  0.0333  0.0333  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  3       24
215     0.2632  -0.2885 0.2415  -0.2438 0.2224  -0.1577 0.5517  0.4853  0.3801  0.2427265910    13      0.52    1       0.2131  4       39                      456460.90
216     0.0855  0.0855  0.1873  0.1873  -0.0596 -0.0596 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  3       21      1       22      213135.15
217     0.1574  0.0746  0.0961  0.0534  0.0464  -0.1717 0.0828  0.0427  0.2181  0.0018391624    2       0.10    0.1     1.0000  4       3                       348582.22
218     0.1546  -0.4238 0.0834  -0.3640 0.1463  -0.3766 0.5784  0.4474  0.5229  0.3227372673    11      2.36    4       0.2444  3       34      9       43      434588.06
219     0.2632  -0.4238 0.2415  -0.4377 0.2224  -0.3766 0.6870  0.6792  0.5990  0.6666339819    40      1.24    3       0.5000  3       36      5       41      528343.33
220     0.3395  -0.4238 0.2415  -0.1881 0.2224  -0.1832 0.7633  0.4296  0.4056  0.3172222878    40      1.95    4       0.4938  3       38      8       46      576279.75
221     0.2130  0.1439  0.2105  0.0023  -0.1115 -0.1758 0.0691  0.2082  0.0643  0.0022063579    2       0.55    1       0.4000  3       32                      254877.10
222     0.2632  0.0855  0.2415  -0.0501 -0.0092 -0.1717 0.1777  0.2916  0.1625  0.0200832745    5       0.64    1       0.3125  4       35                      351813.56
223     0.0806  0.0806  0.1301  0.1301  0.0029  0.0029  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  2       31                      84079.08
224     -0.2247 -0.2885 0.0514  -0.1361 0.0333  -0.0690 0.0638  0.1875  0.1023  0.0029187964    2       0.10    0.1     0.4000  4       16                      451705.73
225     -0.2247 -0.4238 0.0401  -0.1361 0.0626  -0.0690 0.1991  0.1762  0.1316  0.0110113170    3       1.03    2       0.6000  3       31      9       40      577909.94
226     0.1751  0.1751  -0.1003 -0.1003 0.0817  0.0817  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  1       37                      550290.74
227     0.2377  0.0450  0.0869  -0.1881 0.2224  -0.1797 0.1927  0.2750  0.4021  0.0508223442    6       0.85    1       0.2609  1       25                      550290.74
228     0.2283  0.1300  0.2105  -0.1015 -0.0675 -0.1727 0.0983  0.3120  0.1052  0.0076953799    4       0.55    1       0.4444  1       34                      269438.24
229     0.2130  0.2130  0.2105  0.2105  -0.1115 -0.1115 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  1       36                      196970.98
230     0.1651  0.0790  0.0981  -0.2438 0.1557  -0.0284 0.0861  0.3419  0.1841  0.0129259435    3       1.33    2       1.0000  1       32                      112616.71
231     0.4230  0.2283  0.0714  -0.1848 0.2484  -0.1084 0.1947  0.2562  0.3568  0.0424498475    4       0.78    1       0.4444  1       36                      185393.37
232     -0.1432 -0.1432 0.1086  0.1086  0.0062  0.0062  0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  3       27      1       28      29255.65
233     0.2130  0.2130  0.2105  0.2105  -0.1115 -0.1115 0.0000  0.0000  0.0000  0.0000000000    1       1.00    1       1.0000  3       34      1       35      145316.22
234     0.2283  0.2127  0.0636  -0.0501 -0.1084 -0.1132 0.0156  0.1137  0.0048  0.0000203064    2       0.10    0.1     1.0000  2       32      7       39      33202.35
235     0.3348  0.1685  -0.0166 -0.1848 0.2484  0.0525  0.1663  0.1682  0.1959  0.0130694974    5       1.40    3       0.7143  1       28      1       29      118646.24
236     0.1574  0.1574  0.0961  0.0961  -0.1717 -0.1717 0.0000  0.0000  0.0000  0.0000000000    1       0.10    0.1     1.0000  3       31      3       34      242306.02
237     0.3388  0.0044  0.2415  -0.1003 0.1578  -0.0284 0.3344  0.3418  0.1862  0.0507603041    9       0.90    1       0.5294  1       7       0       7       152392.35

Appendix 3. Environmental variables measured and ordination axis positions of the 92 plots. Abbreviations: Marg., marginal ridge; Pied., piedmont; Flat, flat highlands; Valley, secondary valley; Bott., bottomhill; Subw., subwet; Sat., saturated; Conc., concave endorreic; F. end., flat endorreic; Drain., drained; Bil.sl., bilateral symmetry in the slope direction; Bil.or., bilateral symmetry orthogonal to slope direction; Ellip., ellipsoid; Heter., heterometric radial symmetry; Water, water body; Grass, grassland; Sedge, sedge-rushland; Shrub, shrubland; M. Sav., man-made savanna; E. oaks, evergreen oaks; D. oaks, deciduous oaks; Pine, pine forest; Alpine, alpine shrubland; Veg. alt., vegetation altitudinal belt; Soil W., soil wetness; Cond., conductivity (microsiemens/cm); concentrations of Na, K, Ca, Mg, Cl, SO4, CO3H, NO3, and P are in meq/L; O.M., organic matter; W. index, water index.

A tab-delimited version of this appendix is available as a text file.

      Morphotectonic units             Geomorphological units        Slope     Topographic position       Soil wetness         External drainage            Morphometry (shape)                                "Plot ""vegetation"" type"                           Veg. alt.                                                                                                                                                                        Ordination axis coordinates
Plot  Ridge   Marg.   Pied.   Slope   Flat  Valley    Basin   Pied. Slope>4o  Bott.   Slope   Uphill   Subw.   Wet    Sat.    Conc.  F.end.  Drain.  Bil.sl. Bil.or.  Ellip.   Heter.  Water   Grass  Sedge   Shrub   M.Sav.  Forest  E.oaks  D.oaks   Pine  Alpine  Alt.(m) Slope(o)   Soil W.  pH    Cond.       Na      K     Ca        Mg      Cl      SO4    CO3H     NO3     P   O.M.(%)    C      N(%)    AET   W.index Sand(%) Clay(%) Silt(%)   Axis 1  Axis 2  Axis 3
1       0       0       1       1       0       0       0       0       0       1       0       0       1       0       0       0       0       1       1       0       0       0       0       0       0       0       1       0       1       0       0       0       560     1       -1.83   6.7     72      0.17391 0.10256 0.25    0.10833 0.07887 0.11042 0.51967 0.02097         2.11    1.23    0.114   414.3   -8.3    76      10      14      -0.2362 0.0186  -0.1247
2       0       0       1       0       0       1       0       0       0       1       0       0       1       0       0       0       1       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       720     1       -1.209  6.6     63.4    0.30435 0.07692 0.15    0.08333 0.11549 0.15    0.31148 0.07581 0.18868 4.02    2.34    0.173   404     21.8    66      10      24      -0.2314 -0.0740 -0.0628
3       0       0       1       0       0       1       0       0       0       1       0       0       1       0       0       0       1       0       0       0       1       0       0       0       1       0       0       0       1       0       0       0       720     1       -1.209  6.3     93.1    0.34783 0.0641  0.3     0.11667 0.26197 0.2625  0.15574 0.12258 0.16038 4.22    2.45    0.185   404     21.8    64      12      24      -0.2308 -0.1706 -0.1177
4       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       0       1       1       0       0       0       0       1       0       0       0       0       1       0       0       0       700     1       -1.209  6.7     59.5    0.21739 0.03846 0.25    0.06667 0.11549 0.10833 0.31148 0.03065 0.26415 3.45    2.01    0.153   416.4   -8.3    66      12      22      -0.2843 0.0401  -0.0682
5       0       0       1       0       0       0       0       1       0       0       1       0       0       1       0       0       1       0       0       1       0       0       0       0       0       0       1       0       1       0       0       0       900     1       -1.519  6.9     58.5    0.13043 0.15385 0.2     0.09167 0.09859 0.07708 0.15574 0.17581 0.37736 6       3.49    0.273   448.7   28.5    57      12      31      -0.2885 0.0514  0.0333
6       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       0       0       0       1       0       1       0       0       0       0       1       0       0       0       900     0       -1.103  6.8     78.9    0.30435 0.07692 0.25    0.125   0.14648 0.11042 0.41639 0.07097 0.14151 5.06    2.94    0.207   448.7   28.5    40      20      40      -0.3396 -0.0592 0.0704
7       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       0       1       0       1       0       0       0       900     0       -1.103  6.6     60.1    0.17391 0.07692 0.3     0.03333 0.09859 0.12083 0.41639 0.00806 0.17925 5.09    2.96    0.221   426.1   46.3    67      13      20      -0.2312 -0.0008 -0.0073
8       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       0       0       1       0       0       0       0       0       0       1       1       0       0       0       900     0       0.801   6.18    94.2    0.34783 0.03846 0.325   0.08333 0.11549 0.1375  0.15574 0.45806 0.19811 4.52    2.63    0.194   437.3   3.1     60      9       31      -0.2247 -0.1361 -0.0690
9       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       0       0       1       0       0       0       0       0       1       0       1       0       0       0       900     0       -1.519  6.19    70.8    0.26087 0.02564 0.25    0.08333 0.11549 0.11875 0.20656 0.22258 0.16038 5.93    3.45    0.26    437.3   3.1     36      16      48      -0.3526 -0.0961 0.0883
10      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       1       0       0       0       1       0       0       0       0       1       0       0       0       1120    4.5     0.623   6.76    65      0.21739 0.02564 0.25    0.05833 0.11268 0.11042 0.31148 0.07419 0.13208 4.15    2.42    0.168   417.9   117.6   66      11      23      0.0452  0.0608  -0.0709
11      1       0       0       1       0       0       0       0       0       0       1       0       0       1       0       0       0       1       0       1       0       0       0       1       0       0       0       0       1       0       0       0       1120    2       0.478   6.36    63.2    0.26087 0.03846 0.15    0.06667 0.13239 0.12917 0.15574 0.16774 0.13208 4.72    2.75    0.221   417.9   117.6   68      7       25      0.0455  -0.0108 -0.0822
12      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       1       0       0       0       0       0       0       1       0       0       0       1       0       0       1140    4.5     0.317   6.39    84.1    0.26087 0.03846 0.3     0.1     0.12113 0.13542 0.36393 0.36774 0.12264 4.32    2.51    0.216   418.9   114.1   60      10      30      0.0214  -0.0047 -0.0916
13      1       0       0       1       0       0       0       0       0       0       1       0       0       0       1       0       0       1       0       1       0       0       0       1       0       0       0       0       0       1       0       0       1240    5       0.822   6.04    127.8   0.30435 0.12821 0.4     0.2     0.20282 0.19792 0.31148 0.37581 0.15094 7.54    4.38    0.341   409.2   143.6   68      7       25      0.1300  -0.1015 -0.1727
14      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       1       0       0       1240    5       0.822   6.32    81      0.26087 0.05128 0.25    0.14167 0.18592 0.20625 0.15574 0.17419 0.10377 4.69    2.73    0.177   409.2   143.6   72      8       20      0.1251  -0.0358 -0.1467
15      0       0       1       0       0       0       0       1       1       1       0       0       0       1       0       0       0       0       1       0       1       0       0       0       0       0       1       0       1       0       0       0       1000    1       0.444   5.85    165.6   0.26087 0.17949 0.65    0.31667 0.14366 0.30625 0.20656 0.87097 0.39623 9.08    5.28    0.444   429.5   66.6    74      6       20      -0.0965 -0.1800 -0.2050
16      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       1       0       0       0       0       0       1       0       0       1       0       0       0       1250    7       0.801   6.23    56.6    0.21739 0.03846 0.175   0.05    0.15211 0.42292 0.20656 0.07258 0.09434 4.09    2.38    0.189   408.4   144.2   76      5       19      0.1496  0.0078  -0.1388
17      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       1       0       0       0       0       0       1       0       0       1       0       0       0       1250    19      0.516   5.86    52.3    0.17391 0.03846 0.15    0.05    0.14366 0.06875 0.20656 0.11774 0.09434 5.23    3.04    0.221   408.4   144.2   73      5       22      0.2037  0.1094  -0.1120
18      1       0       0       0       0       1       0       0       0       1       0       0       1       0       0       1       0       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       1165    0       0.19    5.7     58.6    0.19565 0.03846 0.2     0.05833 0.16338 0.09375 0.31148 0.13871 0.22642 1.57    0.92    0.067   416.5   114.5   67      11      22      -0.0244 -0.1487 0.0187
19      1       0       0       0       1       0       0       0       0       1       0       0       0       0       1       0       1       0       0       0       0       1       0       0       0       0       0       1       0       0       1       0       1560    0       0.842   5.78    93.6    0.26087 0.12821 0.225   0.09167 0.18592 0.30625 0.31148 0.09839 0.23585 12.4    7.21    0.585   431.8   134.9   74      3       23      0.1447  -0.2438 0.0446
20      1       0       0       0       1       0       0       0       0       0       0       1       0       0       1       0       1       0       1       0       0       0       0       0       0       0       1       0       0       0       1       0       1560    0       0.822   5.56    101.7   0.52174 0.05128 0.2     0.11667 0.3493  0.16667 0.20656 0.12258 0.12264 12.3    7.11    0.563   431.8   134.9   53      26      21      0.1620  -0.4377 0.1503
21      0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       890     1       -1.103  6.45    178.4   0.91304 0.23077 0.45    0.59167 0.47606 0.44167 0.62459 0.03065 0.18868 6.37    3.7     0.264   427.1   -5.3    72      5       23      -0.2724 -0.2719 -0.3119
22      0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       890     0       -1.103  6.5     284     1.08696 0.10256 0.95    0.51667 0.64789 0.75    0.41639 0.15806 0.16038 5.49    3.19    0.2     427.1   -5.3    68      12      20      -0.2992 -0.3640 -0.3766
23      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       1       0       0       0       0       0       0       1       0       0       0       1       0       1490    10      0.822   5.95    99.7    0.26087 0.28205 0.35    0.10833 0.19437 0.2     0.51967 0.05323 0.17925 14.61   8.47    0.34    391.6   119.6   72      9       19      0.2127  -0.0501 -0.1132
24      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       0       0       0       1       0       0       0       1       0       1480    14      0.822   6.13    144.1   0.43478 0.07692 0.55    0.15833 0.2507  0.3             0.55645 0.09434 6.5     3.78    0.163   391.6   119.6   68      10      22      0.2283  0.0636  -0.1084
25      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       1       0       0       0       1195    15      0.801   6.21    108.7   0.47826 0.07692 0.35    0.13333 0.23099 0.2625  0.51967 0.03871 0.16038 10.72   6.23    0.291   416.3   114.4   71      8       21      0.1439  0.0023  -0.1758
26      1       0       0       1       0       0       0       0       0       1       0       0       1       0       0       0       1       0       1       0       0       0       0       1       0       0       0       0       1       0       0       0       1185    0       -1.83   6.19    72.2    0.26087 0.05128 0.25    0.08333 0.19718 0.19375 0.2082  0.10806 0.17925 3.22    1.87    0.098   416.3   114.4   81      1       18      -0.0818 -0.0733 -0.0647
27      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       1       0       0       0       1205    16      0.643   6.16    63.7    0.21739 0.12821 0.2     0.06667 0.16056 0.13125 0.2082  0.07581 0.11321 3.89    2.26    0.11    416.3   114.4   67      8       25      0.1574  0.0961  -0.1717
28      1       0       0       1       0       0       0       0       1       0       0       1       1       0       0       0       1       0       0       1       0       0       0       0       0       0       0       1       0       0       1       0       1260    5       -0.404  6.26    105.3   0.26087 0.07692 0.45    0.125   0.11268 0.12292 0.31148 0.47581 0.32075 7.64    4.44    0.304   447.3   67.6                            -0.0197 -0.0945 0.0016
29      1       0       0       1       0       0       0       0       0       0       1       0       0       0       1       0       1       0       0       1       0       0       0       1       0       0       0       0       0       0       1       0       1185    4       0.842   5.83    86.3    0.34783 0.10256 0.2     0.08333 0.12676 0.38333 0.41639 0.1     0.09434 19.1    14.1    0.645   423.5   103.6   64      8       28      0.1054  -0.1288 -0.0372
30      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       0       1       0       1180    8       0.842   5.87    78.3    0.30435 0.20513 0.25    0.08333 0.14366 0.2375  0.31148 0.07097 0.13208 8.98    5.22    0.304   423.5   103.6   71      7       22      0.0958  -0.0330 -0.0989
31      1       0       0       1       0       0       0       0       1       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       0       0       0       0       0       1       0       1145    5       0.801   6.1     55.9    0.21739 0.05128 0.2     0.08333 0.08169 0.10417 0.51967 0.00323 0.09434 6.5     3.78    0.233   422.5   105     69      5       26      0.0450  0.0029  -0.0410
32      1       0       0       0       0       1       0       0       0       0       1       0       0       0       1       0       1       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       1340    3       0.801   5.87    91.3    0.43478 0.10256 0.2     0.05    0.16338 0.22292 0.31148 0.20645 0.08491 11.86   6.89    0.47    400.1   142.9   69      5       26      0.1474  -0.2032 -0.0702
33      1       0       0       0       0       1       0       0       1       0       1       0       0       1       0       0       1       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       1340    6       -0.998  6.57    32.3    0.08696 0.05128 0.025   0.025   0.08732 0.05833 0.36393 0.04355 0.06604 8.91    5.18    0.291   400.1   142.9   77      3       20      0.0746  0.0534  0.0464
34      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       1       0       0       0       0       0       0       0       1       0       0       1       0       1390    9       0.822   6.63    46.6    0.15217 0.05128 0.05    0.025   0.10986 0.09375 0.41639 0.04032 0.13208 14.47   8.41    0.452   396.3   155.2   68      6       26      0.2377  0.0869  -0.0481
35      1       0       0       1       0       0       0       0       0       1       0       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       1       0       0       1300    3       0.842   6.62    71.3    0.19565 0.07692 0.15    0.05    0.10986 0.29167 0.31148 0.21935 0.12264 13.33   7.75    0.445   404.8   139.6   63      3       34      0.1302  -0.0531 -0.0419
36      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       1       0       0       0       1       0       0       0       0       0       0       1       0       1550    13      0.444   6.29    75.2    0.21739 0.11538 0.125   0.05833 0.34366 0.07708 0.46721 0.00806 0.15094 12.73   7.4     0.417   405.1   157.9   71      3       26      0.2236  0.0174  -0.0517
37      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       0       0       0       0       1       0       0       1       0       1550    13      0.822   6.94    56.9    0.17391 0.05128 0.15    0.05833 0.19718 0.06875 0.63279 0.01613 0.09434 7.44    4.32    0.256   405.1   157.9   68      6       26      0.2174  0.1255  -0.0675
38      1       0       0       1       0       0       0       0       1       1       0       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       1       0       0       1400    12      0.801   6.8     94.8    0.43478 0.02564 0.2     0.09167 0.50141 0.2125  0.26066 0.06129 0.27358 5.03    2.92    0.203   408.4   153.2                           0.1546  -0.0441 -0.1577
39      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       1       0       0       0       1       0       0       0       0       0       1       0       0       1390    8       0.842   6.38    141.8   0.34783 0.14103 0.45    0.20833 0.62817 0.46042 0.31148 0.10645 0.10377 16.42   9.54    0.755   408.6   153.3   82      2       16      0.2010  -0.1812 -0.1797
40      1       0       0       0       0       0       1       0       1       1       0       0       0       0       1       0       0       1       1       0       0       0       0       0       0       0       0       1       0       1       0       0       1250    13      0.822   6.82    40.5    0.13043 0.05128 0.1     0.04167 0.0338  0.15    0.31148 0.05323 0.15094 10.45   6.08    0.266   443.9   110.4   73      2       25      0.0855  0.1873  -0.0596
41      1       0       0       0       0       0       1       0       1       0       1       0       0       1       0       0       0       1       0       1       0       0       0       0       0       0       1       0       0       1       0       0       1280    11      -1.209  6.8     44.6    0.13043 0.07692 0.1     0.025   0.07606 0.06458 0.26066 0.11129 0.12264 6.57    3.82    0.284   430.9   131.9   78      2       20      0.0249  0.1878  -0.0477
42      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       0       1       0       0       0       0       0       1       0       1500    8       -1.209  6.92    42.9    0.08696 0.02564 0.05    0.00833 0.07887 0.075   0.31148 0.12419 0.06604 21.51   12.5    0.83    437.5   123.8   75      8       17      0.1198  0.1438  0.0662
43      1       0       0       0       1       0       0       0       0       1       0       0       0       0       1       1       0       0       0       0       1       0       1       0       0       0       0       0       0       0       1       0       1490    0       0.842   6.63    33.3    0.08696 0.05128 0.1     0.03333 0.03662 0.04375 0.2082  0.1     0.16981 5.49    3.19    0.204   436.6   125.9   66      6       28      0.0790  -0.0651 0.1557
44      0       1       0       1       0       0       0       0       1       0       1       0       1       0       0       0       0       1       0       1       0       0       0       0       0       1       0       0       1       0       0       0       970     15      -1.3    6.64    26.6    0.04348 0.02564 0.05    0.00833 0.05915 0.03958 0.26066 0.0371  0.11321 3.15    1.83    0.093   457.5   9       56      15      29      -0.1727 0.3545  -0.0643
45      0       0       1       0       0       0       0       1       1       0       1       0       1       0       0       0       0       1       0       0       1       0       0       1       0       0       0       0       1       0       0       0       980     5       -0.998  6.62    67.1    0.21739 0.10256 0.05    0.08333 0.13803 0.11667 0.26066 0.16774 0.09434 5.96    3.47    0.242   458.9   9.2     62      10      28      -0.2362 0.0593  -0.0049
46      0       0       1       0       0       0       0       1       1       0       1       0       1       0       0       0       0       1       0       1       0       0       0       0       1       0       0       0       1       0       0       0       840     6       -1.83   6.95    20.8    0.04348 0.02564 0.05    0.01667 0.03099 0.02917 0.2082  0.01129 0.11321 2.51    1.46    0.091   458.9   12.7    55      20      25      -0.2838 0.2423  0.0702
47      0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       0       0       0       1       0       1       0       0       0       0       1       0       0       0       1000    2       -1.103  6.83    39.5    0.06522 0.0641  0.025   0.06667 0.10141 0.0625  0.2082  0.05323 0.12264 5.83    3.39    0.168   483.9   10.1    64      12      24      -0.3387 0.0733  0.1412
48      0       0       1       0       0       0       0       1       1       1       0       0       0       1       0       0       0       1       0       1       0       0       0       1       0       0       0       0       1       0       0       0       935     12      -1.519  6.62    35.9    0.08696 0.03846 0.075   0.03333 0.09296 0.05625 0.41639 0.09677 0.12264 2.98    1.73    0.165   471.7   5.8     64      10      26      -0.2593 0.2071  0.0364
49      0       0       1       0       0       0       0       1       1       1       0       0       1       0       0       0       0       1       0       0       1       0       0       1       0       0       0       0       1       0       0       0       935     5       -1.83   6.65    25.5    0.04348 0.02564 0.05    0.00833 0.05634 0.03958         0.05968 0.14151 2.99    1.74    0.124   471.7   5.8                             -0.2741 0.0585  0.0158
50      1       0       0       0       1       0       0       0       0       0       1       0       0       0       1       0       1       0       0       1       0       0       0       1       0       0       0       0       0       0       0       1       1680    3       0.822   6.64    62      0.1087  0.07692 0.075   0.01667 0.10423 0.09792 0.31148 0.14677         23.13   13.44   0.945   424     148.9   72      3       25      0.1685  -0.1236 0.0894
51      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       0       1       0       1680    14      0.096   6.98    44.6    0.13043 0.05128 0.1     0.025   0.12394 0.12708 0.26066 0.09032 0.23585 11.56   6.72    0.365   417.8   150.7   66      7       27      0.2101  0.1602  -0.0017
52      1       0       0       0       1       0       0       0       1       1       0       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       0       0       1       1860    16      0.801   6.86    41.5    0.04348 0.10256 0.05    0.00833 0.10986 0.06458 0.31148 0.01613 0.23585 16.49   9.59    0.875   388.8   150.9                           0.3395  0.0536  0.1278
53      1       0       0       0       1       0       0       0       1       1       0       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       0       0       1       1860    9       0.822   6.63    47.8    0.08696 0.12821 0       0.00833 0.09859 0.13333 0.26066 0.06129 0.31132 25.25   14.68   0.875   388.8   150.9   84      3       13      0.3305  -0.0456 0.1578
54      1       0       0       1       0       0       0       0       1       1       0       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       0       0       1       1850    18      0.516   6.54    42.5    0.08696 0.05128 0.05    0.00833 0.10423 0.17083 0.36393 0.04194 0.16981 23.4    7.79    0.682   399     143.6                           0.3388  0.1321  -0.0089
55      1       0       0       1       0       0       0       0       1       0       0       1       0       0       1       0       0       1       0       1       0       0       0       1       0       0       0       0       0       0       0       1       1860    6       0.842   6.11    59      0.13043 0.41026 0.025   0.00833 0.22254 0.0625  0.41639 0.00323         58.31   33.9    2.21    392.7   148.4   80      2       18      0.2757  -0.1036 0.0525
56      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       0       0       0       0       1       0       0       1       0       1520    15      0.822   6.26    21      0.04348 0.02564 0.025   0.03333 0.07887 0.03958 0.15574 0.01129 1.18868 3.99    2.32    0.63    395.5   142.3   59      6       35      0.2632  0.2415  -0.0092
57      1       0       0       0       1       0       0       0       1       0       1       0       0       0       1       0       0       1       0       1       0       0       0       0       0       0       0       1       0       0       1       0       1590    11      0.801   6.64    61.4    0.08696 0.05128 0.45    0.01667 0.09296 0.125   0.62459 0.04355 0.5     12.71   7.39    0.442   412.6   160     48      16      36      0.2174  0.0724  0.0935
58      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       0       1       0       0       0       0       0       1       0       0       1       0       0       1340    10      -1.103  6.2     75.1    0.13043 0.17949 0.15    0.175   0.11268 0.09167 0.2082  0.46613 0.13208 8.93    5.19    0.442   475.7   99.1    66      4       30      -0.0615 0.0531  0.0583
59      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       1       0       0       0       0       0       0       0       1       0       0       1       0       1610    12      0.842   6.15    70.7    0.21739 0.05128 0.05    0.05    0.17465 0.22708 0.2082  0.12581 0.17925 22.33   12.98   0.696   470.9   128.7   54      8       38      0.1773  0.0348  0.0620
60      1       0       0       0       0       0       1       0       1       0       1       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       0       0       1       0       0       0       1220    13      -0.509  5.95    31      0.08696 0.05128 0.025   0.025   0.10986 0.07708 0.31148 0.02258 0.10377 5.85    3.4     0.254   466.9   58      66      6       28      -0.0791 0.1838  0.0573
61      1       0       0       0       1       0       0       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       0       0       1       0       0       0       0       0       1       0       1460    2       0.623   6.65    54      0.15217 0.02564 0.25    0.03333 0.13521 0.0875  0.41639 0.03387 0.08491 13.74   7.99    0.564   435     134.5   87      1       12      0.1579  -0.1881 0.2224
62      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       1       0       0       0       0       0       0       0       1       0       0       1       0       1460    16      0.822   6.94    86.3    0.13043 0.07692 0.95    0.06667 0.09014 0.18542 1.24754 0.03226 0.21698 19.76   11.49   0.31    435     132.8                           0.2130  0.2105  -0.1115
63      1       0       0       1       0       0       0       0       0       0       1       0       0       0       1       1       0       0       1       0       0       0       1       0       0       0       0       0       0       0       1       0       1280    0       0.842   6.79    357     0.23913 0.4359  3       0.79167 0.15775 1.06042 1.27213 0.22581 0.78302 33.71   19.6    1.13    449     115     78      2       20      0.0278  0.1327  0.1452
64      1       0       0       0       0       0       1       0       0       1       0       0       0       1       0       0       1       0       0       0       1       0       0       0       0       0       0       1       0       1       0       0       1160    0       0.842   6.84    64.6    0.04348 0.05128 0.3     0.15833 0.0507  0.24375 0.31148 0.07581 0.68868 7.22    9.2     0.291   486.6   42      54      8       38      -0.1432 0.1086  0.0062
65      0       0       1       1       0       0       0       0       1       0       1       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       0       0       0       1       0       0       925     6       -1.519  7.95    45.2    0.13043 0.02564 0.125   0.01667 0.11268 0.10625 0.21967         0.07547 4.5     2.62    0.182   467.1   46.6    43      23      34      -0.1999 0.0825  0.0232
66      0       0       1       1       0       0       0       0       0       1       0       0       1       0       0       0       1       0       0       0       0       1       0       1       0       0       0       0       0       1       0       0       920     1       -1.209  7.3     43.4    0.17391 0.02564 0.15    0.025   0.07887 0.11042 0.21967 0.01613 0.08491 4.8     2.79    0.235   467.1   46.6    58      13      29      -0.3087 0.0686  0.1463
67      0       0       1       0       0       0       1       0       0       1       0       0       1       0       0       0       1       0       0       0       0       1       0       1       0       0       0       0       1       0       0       0       830     0       -1.103  7.15    52.4    0.13043 0.01282 0.3     0.025   0.10986 0.19792 0.2082  0.03548 0.12264 4.33    2.52    0.206   465.9   12.1    41      23      36      -0.3533 0.0834  0.0187
68      0       0       1       0       0       0       1       0       0       1       0       0       1       0       0       1       0       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       840     0       -1.83   6.88    62.4    0.15217 0.10256 0.25    0.04167 0.09014 0.15208 0.1541  0.09032 0.75472 7.57    4.4     0.413   466.3   15      28      38      34      -0.3924 0.0560  0.1324
69      0       0       1       0       0       0       0       1       0       1       0       0       0       0       1       1       0       0       0       0       1       0       1       0       0       0       0       0       1       0       0       0       1030    0       0.842   6.86    56.8    0.17391 0.01282 0.275   0.00833 0.08451 0.16667 0.21967         0.12264 8.73    5.07    0.446   447.4   93.7    28      38      34      -0.1845 -0.1419 0.1870
70      0       0       1       0       0       0       0       1       0       1       0       0       0       0       1       1       0       0       0       0       1       0       1       0       0       0       0       0       1       0       0       0       1030    0       0.842   6.86    56.8    0.17391 0.01282 0.275   0.00833 0.08451 0.16667 0.21967         0.12264 8.73    5.07    0.446   447.4   93.7    79      6       15      -0.1845 -0.1419 0.1870
71      0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       1       0       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       1040    0       0.444   6.87    32.9    0.1087  0.01282 0.1     0.00833 0.07324 0.10208 0.1541          0.07547 2.88    1.67    0.125   447.4   93.7    29      21      50      -0.0955 -0.0792 -0.0060
72      0       0       1       0       0       0       1       0       0       1       0       0       0       1       0       0       1       0       1       0       0       0       0       1       0       0       0       0       1       0       0       0       820     0       -1.519  6.63    85.4    0.21739 0.02564 0.425   0.125   0.05352 0.40625 0.28525 0.03871 0.17925 3.62    2.1     0.179   456.6   3.2     65      3       32      -0.4238 -0.0139 0.0626
73      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       1       0       0       0       0       0       0       1       0       0       1       0       0       1480    12      -0.382  6.48    53.7    0.1087  0.02564 0.25    0.025   0.08169 0.12917 0.21967 0.02581 0.10377 8.66    5.03    0.417   420     73.1    61      3       36      0.0806  0.1301  0.0029
74      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       1       0       0       0       0       0       0       1       0       0       1       0       0       1460    6       -0.204  6.46    40.6    0.08696 0.02564 0.15    0.01667 0.09296 0.11667 0.13279 0.0129  0.31132 6.63    3.86    0.326   420     73.1    69      2       29      0.0395  0.1106  0.0444
75      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       1       0       0       0       0       0       0       1       0       0       0       0       0       1       1450    20      0.696   6.11    55.9    0.1087  0.02564 0.2     0.025   0.22535 0.11667 0.16557 0.00645 0.63208 6.77    3.93    0.274   419.8   80.5    70      2       28      0.1645  0.1733  -0.0587
76      1       0       0       1       0       0       0       0       0       1       0       0       0       0       1       0       0       1       1       0       0       0       0       0       1       0       0       0       0       0       0       1       1520    1       0.822   5.8     65.8    0.13043 0.0641  0.2     0.04167 0.12113 0.18542 0.21967 0.00323 0.12264 27.92   16.23   1.02    421.6   92      68      4       28      0.1751  -0.1003 0.0817
77      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       1       0       0       0       0       0       0       0       1       0       0       1       0       0       1330    16      0.842   5.77    58.4    0.1087  0.0641  0.2     0.04167 0.08451 0.13542 0.21967 0.00323 0.12264 11.77   6.85    0.48    424.4   85.4    40      13      47      0.1651  0.0981  -0.0284
78      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       1       0       0       0       0       0       0       1       0       0       0       1       0       0       1230    16      0.623   6.04    31.3    0.1087  0.01282 0.1     0.01667 0.05634 0.08333 0.19836 0.00323 0.07547 3.47    2.02    0.125   450.4   77.9    57      14      29      0.0044  0.1602  0.1070
79      1       0       0       0       0       0       0       1       0       0       1       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       0       0       1       0       0       0       1060    1       -1.209  6       34.3    0.08696 0.02564 0.125   0.01667 0.0507  0.09375 0.21967         0.14151 3.9     2.27    0.187   464.5   35.5    68      13      19      -0.1964 0.0221  0.0441
80      1       0       0       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       1       0       0       0       0       0       1       0       0       0       0       1       0       0       1070    1       -0.998  5.96    37.7    0.13043 0.02564 0.15    0.025   0.06479 0.11042 0.23115         0.14151 3.74    2.12    0.166   472.8   36      71      8       21      -0.2098 -0.0231 0.0463
81      0       1       0       0       0       0       1       0       0       0       1       0       0       1       0       0       1       0       0       0       0       1       0       0       0       0       1       0       1       0       0       0       910     0       0.444   5.87    55.6    0.17391 0.01282 0.25    0.06667 0.08169 0.15208 0.24098         0.09434 6.56    3.82    0.283   456.4   29      77      2       21      -0.1864 0.1009  -0.1779
82      0       1       0       0       0       0       1       0       0       1       0       0       0       0       1       0       1       0       1       0       0       0       0       1       0       0       0       0       1       0       0       0       920     1       0.822   5.72    78.7    0.23913 0.02564 0.35    0.09167 0.10704 0.23958 0.31967         0.11321 18.18   10.57   0.778   456.4   29      71      3       26      -0.1722 0.0788  -0.1832
83      1       0       0       0       0       0       0       1       1       1       0       0       0       1       0       0       0       1       1       0       0       0       0       0       0       0       1       0       0       1       0       0       1050    5       0.317   5.79    67.6    0.13043 0.07692 0.3     0.025   0.11268 0.11042 0.21967 0.15484 0.16038 3.43    1.99    0.136   450.2   52.9    39      28      33      -0.0813 0.0143  -0.0425
84      0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       1       0       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       920     0       -1.83   5.94    46.5    0.13043 0.05128 0.175   0.03333 0.05634 0.12917 0.19836 0.00323 0.26415 4.73    2.75    0.254   447     9.5     31      32      37      -0.3544 -0.0385 0.1802
85      0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       0       0       0       1       0       0       0       0       1       0       1       0       0       0       960     0       -1.83   5.82    49.3    0.19565 0.02564 0.15    0.025   0.06197 0.17083 0.1541  0.00323 0.32075 11.36   6.61    0.341   457.5   38.7    49      20      31      -0.3323 -0.0579 0.2181
86      0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       0       1       1       0       0       0       0       0       0       0       1       0       1       0       0       0       940     1       -1.103  5.85    53.5    0.17391 0.02564 0.225   0.03333 0.11268 0.13958 0.16557 0.00323 0.12264 7.24    4.21    0.354   462.3   39.8    62      3       35      -0.2659 -0.0229 0.1041
87      1       0       0       0       1       0       0       0       0       1       0       0       0       0       1       1       0       0       1       0       0       0       0       1       0       0       0       0       0       0       0       1       2110    4       0.822   5.65    41.5    0.13043 0.05128 0.15    0.025   0.09859 0.12083 0.18689 0.00323 0.13208 15.74   9.15    0.54    421.8   157.9   76      3       21      0.2588  -0.1848 0.2484
88      1       0       0       0       1       0       0       0       1       0       1       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       0       0       1       2060    12      0.822   5.63    64      0.17391 0.05128 0.25    0.04167 0.12958 0.17083 0.21967 0.00323 0.43396 11.64   8.51    0.48    421.8   157.9                           0.3348  -0.0166 0.0564
89      1       0       0       0       1       0       0       0       1       1       0       0       0       0       1       0       0       1       1       0       0       0       0       1       0       0       0       0       0       0       0       1       2040    8       0.842   5.71    47.6    0.13043 0.0641  0.15    0.01667 0.05915 0.075   0.16557 0.00323 0.15094 23.74   13.8    0.508   421.8   157.9                           0.3094  -0.0788 0.1950
90      1       0       0       1       0       0       0       0       1       0       1       0       0       0       1       0       0       1       0       0       0       1       0       1       0       0       0       0       0       0       0       1       1960    26      0.842   5.49    77.5    0.21739 0.15385 0.2     0.04167 0.07324 0.13958 0.23115 0.00323 0.24528 26.6    15.46   0.95    417.7   160                             0.4230  0.0714  0.0501
91      1       0       0       0       1       0       0       0       0       1       0       0       0       0       1       0       0       1       0       0       0       1       0       1       0       0       0       0       0       0       0       1       1940    0       0.842                   0.21739 0.15385 0.2     0.04167 0.07324 0.13958 0.23115 0.00323 0.24528                         417.7   160     66      3       31      0.2947  -0.2314 0.1976
92      1       0       0       1       0       0       0       0       1       0       1       0       0       1       0       0       0       1       0       0       0       1       0       1       0       0       0       0       0       0       1       0       1820    11      0.822   5.32    76.3    0.17391 0.10256 0.25    0.05833 0.07042 0.16667 0.21967 0.00323 0.20755 21.46   12.48   0.844   413.9   159.5                           0.3221  -0.0297 0.0454

Appendix 4 Details of the multivariate methods used to determine the environmental volume of the species.

In each plot, we recorded 53 variables (Appendix 3) that were ultimately reduced into a three-dimensional, multivariate space. This reduction was done through a two-step, multivariate analysis. The first step consisted of reducing the 53 variables to 12 variables, several of which were multivariate composites.

The first set of three composite variables were derived from 32 binary variables consisting of information on land form (type of morphotectonic unit and type of geomorphological unit), topography, drainage, grassland shape, vegetation physiognomy, and altitude zone. Land form variables were inferred from 1:50000 geological maps, and the rest of variables in this group were inferred from photo-interpretation of 1:20000 panchromatic stereopairs. Composite variables were derived from the first three eigenvectors of a correspondence analysis. These three eigenvectors primarily represented landscape position of the wet grasslands.

The next set of three composite variables consisted of the first three eigenvectors of a PCA based on soil chemical composition: pH, conductivity, sodium, potassium, calcium, magnesium, chloride, sulfate, carbonate and bicarbonate, organic matter, nitrate, nitrogen, and phosphorous. We took five 20-cm depth soil samples regularly distributed within each plot. These five samples were mixed into one composite sample for analyses of soil chemical composition and soil texture. The following soil chemical composition variables were analyzed in a 1:10 soil solution: pH, conductivity, sodium, potassium, calcium, magnesium, chloride, sulfate, carbonate, bicarbonate, and nitrate. Organic matter, nitrogen, and phosphorous were directly analyzed in the soil sample. These three eigenvectors were interpreted as total ion content, proportion of organic matter, and soil acidity.

The seventh composite variable was soil texture, based on the first eigenvector of a PCA of the percentages of sand, silt, and clay. The other five variables were: altitude, actual evapotranspiration, water index, ground slope, and soil wetness. Altitude was estimated with a calibrated altimeter in the field. Actual evapotranspiration and water index were inferred from interpolation of the region climate stations. Ground slope was estimated with a clinometer in the field. Soil wetness was estimated by means of an index based upon qualitative indicators such as gley traits and percentage of saturation. Across all variables, <1% of the data were missing, primarily texture (11%) and nitrate (9%). In these instances, we substituted the mean value for that variable.

These 12 variables defined an environmental space based on the first three eigenvectors from a PCA. This single space was then used to determine the volumes for all species. The distribution of the sites in the space is shown in Fig. 8. All ordination analyses were done with PC-ORD (McCune and Mefford 1995).


Fig. 8. Distribution of the 92 plots in the first three dimensions of the ordination space defined by the 53 environmental variables.

GIF image file (15 K)

The environmental volume of each plant species was measured by determining the maximum and minimum coordinates on all three axes. To do this, for each species on each axis we determined the sites containing that species with the largest and smallest axis scores. The volume was the rectangular solid containing these points. This measure is equivalent to the range volume of Burgman (1989) except that we did not standardize axis length. For species appearing in only one site, the volume was zero. Changing this volume to an arbitrarily chosen small value (e.g., 10% of the smallest two-site volume), would not change the analyses.


Address of Correspondent:
José M. Rey Benayas
Ecología
Facultad de Ciencias
E-28871 Alcalá de Henares
Universidad de Alcalá, Spain
Phone: 34-1-885-4987
Fax: 34-1-885-5090
gljmrb@geolog.alcala.es

*The copyright to this article passed from the Ecological Society of America to the Resilience Alliance on 1 January 2000.

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