Fig. 1. The Doņana social-ecological system, and the ecodistricts that constitute the Greater Fluvial-Littoral Ecosystem of Doņana. The semi-natural ecodistricts are those ecosystems modified by human use throughout history in a process of co-evolution. Transformed ecodistricts are those that have suffered a great modification (degradation) in the second half of the 20th century, mainly due to the introduction of agriculture, forest plantations, or aquiculture. Sample points are those places where interviews and questionnaires were made. The main use in the white areas is agriculture.