Fig. 5. Effects of a shifting attractor on the CLS of a 20-species system. (A) Temporal dynamics of six of the 20 species. Up to generation 2000, all 20 species coexist and community structure is approximately constant because “open” recruitment enables recruits of all species to establish (with equal likelihood) on small patches on the landscape cleared by a low level of disturbance (see Methods). At generation 2001, the “open”recruitment is “turned off” and species begin to go locally extinct (extinction points are indicated by triangles on the x-axis), which leads to different landscape dynamics. Prediction-r2 spectra are shown for analyses based on series of five maps finishing at generation 2000, 4000, and 12 000 (B–D, respectively), where consecutive maps in each series were separated by 10 time steps. Spectra are based on analysis of densities of two species (Sred and Sblue, shown as bold lines in A). Both species indicate a reduction in the CLS from about 55 length units to about 20 length units. Areas identifying the CLS are shown as black horizontal bars.